Quinoa

Quinoa
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* Quinoa is an invaluable crop, highlighted by the FAO as one of the world’s main crops for future food security * Timely publication – The year 2013 has been declared «The International Year of the Quinoa» (IYQ), recognizing the Andean indigenous peoples, who have maintained, controlled, protected and preserved quinoa as food for present and future generations thanks to their traditional knowledge and practices of living well in harmony with mother earth and nature. * Covers the history, phylogeny and systematics, botany and agrotechnology

Оглавление

Atul Bhargava. Quinoa

QUINOA

Contents

Contributors

Preface

Acknowledgements

1 Introduction

1.1 Underutilized Crops

1.2 Chenopodium as an Underutilized Plant

1.3 Quinoa

1.3.1 Nutritional importance of quinoa

1.3.2 Stress tolerance

1.3.3 Economic uses

1.3.4 Medicinal importance

1.4 Concluding Remarks

References

2 Historical Perspectives and Domestication

2.1 Introduction

2.2 General Process of Plant Domestication

2.3 Importance of the Genus Chenopodium and Domestication of C. quinoa

2.4 Current Insights into the Evolution of Genetic Diversity in Quinoa

2.5 Current Typology to Describe the Diversification and Utilization of the Quinoas in South America

2.5.1 Quinoas of the Altiplano (northern Andean highlands)

2.5.2 Quinoas from the Salares (southern highlands)

2.5.3 Quinoas from Inter Andean Valleys

2.5.4 Quinoas from arid zones and dry conditions (western highlands)

2.5.5 Quinoas from high altitudes and cool climate

2.5.6 Quinoas from the coastal regions and near the sea

2.5.7 Quinoas from jungle and tropical zones

2.5.8 Quinoa from high rainfall and humidity zones

2.5.9 Wild relatives of quinoa

2.6 Concluding Remarks

Acknowledgements

References

3 Distribution. 3.1 Introduction

3.2 The Andean Region and Quinoa

3.3 Quinoa in its Native Region

3.3.1 Bolivia

3.3.2 Peru

3.3.3 Chile

3.3.4 Ecuador

3.3.5 Argentina

3.3.6 Colombia

3.4 The European Experiment

3.5 North America

3.6 Quinoa and Africa

3.7 Experiments in Asia

3.8 Concluding Remarks

References

4 Taxonomy. 4.1 Introduction

4.2 Chenopodiaceae and Amaranthaceae

4.3 Chenopod Classification and Quinoa

4.3.1 Genetic interrelationships based on protein analysis

4.3.2 Molecular markers for phylogenetic analysis

4.4 Concluding Remarks

References

5 Cytology and Genome Size. 5.1 Introduction

5.2 Structural/Physical Organization of Plant Chromosomes

5.3 Cytological Techniques

5.4 Cytological Studies in Quinoa

5.5 Nuclear DNA Content

5.6 Concluding Remarks

References

6 Botany. 6.1 Introduction

6.2 Vegetative Parts

6.2.1 Root system

6.2.2 Stem

6.2.3 Leaves

6.3 Reproductive Parts. 6.3.1 Inflorescence

6.3.2 Flower and floral types

6.3.3 Flower clusters or glomeruli

6.3.4 Fruit

6.4 Concluding Remarks

References

7 Crop Production and Management. 7.1 Introduction

7.2 Land Preparation

7.3 Sowing

7.4 Seeding Rates and Plant Density

7.5 Fertilizer Application

7.6 Water Requirements and Irrigation

7.7 Effect of Temperature and Photoperiod on Quinoa

7.8 Weed Control

7.9 Harvesting

7.10 Postharvest

7.10.1 Predrying and drying

7.10.2 Threshing

7.10.3 Cleaning seeds

7.10.4 Drying

7.10.5 Storage

7.11 Concluding Remarks

References

8 Stress Tolerance. 8.1 Introduction

8.2 Abiotic Stresses

8.3 Quinoa as a Stress-Tolerant Crop

8.3.1 Salt resistance in quinoa

Quinoa as a halophyte

Salinity and its impact on agromorphological traits in quinoa

Salinity stress and dehydrins in quinoa

8.3.2 Drought resistance in quinoa

Deficit irrigation

8.3.3 Cold stress

8.3.4 Other stresses

8.4 Concluding Remarks

References

9 Diseases and Pests. 9.1 Introduction

9.2 Diseases

9.2.1 Bacterial diseases

9.2.2 Fungal diseases. Downy mildew

SYMPTOMS OF DOWNY MILDEW

Damping off

Brown stalk rot

Stem gothic spot

Leaf spot

Grey mould

9.2.3 Viral diseases

9.3 Pests

9.3.1 Insects

9.3.2 Birds

9.3.3 Rodents

9.3.4 Nematodes

9.4 Control

9.5 Concluding Remarks

References

10 Breeding. 10.1 Introduction

10.1.1 Breeding objectives in quinoa

10.2 Outcrossing in Quinoa

10.3 Male Sterility

10.3.1 Male sterility in quinoa

10.4 Breeding in Quinoa

10.4.1 Genetic variability

Genetic variability studies in quinoa

10.4.2 Genetic correlation and path coefficient analysis

Correlation and path studies in quinoa

10.4.3 Genetic diversity

Genetic diversity analysis in quinoa

10.4.4 Hybridization experiments

10.4.5 Genotype × environment interaction (GEI)

10.5 Concluding Remarks

References

11 Molecular Studies

11.1 Introduction

11.2 First Generation of Molecular Studies in Quinoa

11.3 Sequence-Arbitrary Methods. 11.3.1 Random amplified polymorphic DNA (RAPD) and directed amplification of minisatellite-region DNA (DAMD) markers

11.3.2 Amplified fragment length polymorphism (AFLP) markers

11.4 Sequence-Dependent Methods. 11.4.1 Simple-sequence repeat (SSR) markers

11.4.2 Single nucleotide polymorphism (SNP) markers

11.5 Genetic Linkage Maps

11.6 Gene Discovery, Functional Genomics and Non-Coding Genotyping

11.7 Molecular Cytogenetics

11.8 Concluding Remarks

References

12 Chemistry. 12.1 Introduction

12.2 Grain

12.2.1 Nutritional components of quinoa seed. Protein

Carbohydrates

Lipids

Minerals

Vitamins

Fibre

Polyphenols

12.2.2 Antinutritional components of quinoa seed

Phytic acid

Protease inhibitors

Tannins

Saponins

12.3 Leaves

12.4 Concluding Remarks

References

13 Saponins. 13.1 Introduction

13.2 Chemical Structure of Saponins

13.3 Quinoa Saponins

13.4 Biosynthesis of Quinoa Saponins

13.5 Genetic Aspects

13.6 Methods for Studying Saponins in Quinoa

13.7 Removal of Saponins from Quinoa Seeds

13.7.1 Moist method

13.7.2 Dry method

13.7.3 Combined method

13.8 Applications of Quinoa Saponins

13.9 Concluding Remarks

References

14 Transparency from Production to Consumption: New Challenges for the Quinoa Market Chain

14.1 Introduction

14.2 Quinoa: Knowledge-Based Food Consumption in a Modern Society

14.3 Contrasts Between the Chilean Case and Other Latin American Models for the Quinoa Market

14.4 Transparent Markets: A New Model for Quinoa and Other Trade Chains

14.5 Concluding Remarks

Acknowledgements

References

Index

Отрывок из книги

QUINOA

Botany, Production and Uses

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Hammer, K., Heller, J. and Engels, J. (2001) Monograph on underutilized and neglected crops. Genetic Resources and Crop Evolution 48, 3–5.

Hariadi, Y., Marandon, K., Tian, Y., Jacobsen, S.-E. and Shabala, S. (2011) Ionic and osmotic relations in quinoa (Chenopodium quinoa Willd.) plants grown at various salinity levels. Journal of Experimental Botany 62, 185–193.

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