Astrobiology

Astrobiology
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A guide to understanding the formation of life in the Universe The revised and updated second edition of Astrobiology offers an introductory text that explores the structure of living things, the formation of the elements required for life in the Universe, the biological and geological history of the Earth, and the habitability of other planets. Written by a noted expert on the topic, the book examines many of the major conceptual foundations in astrobiology, which cover a diversity of traditional fields including chemistry, biology, geosciences, physics, and astronomy. The book explores many profound questions such as: How did life originate on Earth? How has life persisted on Earth for over three billion years? Is there life elsewhere in the Universe? What is the future of life on Earth? Astrobiology is centered on investigating the past and future of life on Earth by looking beyond Earth to get the answers. Astrobiology links the diverse scientific fields needed to understand life on our own planet and, potentially, life beyond. This new second edition: Expands on information about the nature of astrobiology and why it is useful Contains a new chapter “What is Life?” that explores the history of attempts to understand life Contains 20% more material on the astrobiology of Mars, icy moons, the structure of life, and the habitability of planets New ‘Discussion Boxes’ to stimulate debate and thought about key questions in astrobiology New review and reflection questions for each chapter to aid learning New boxes describing the careers of astrobiologists and how they got into the subject Offers revised and updated information throughout to reflect the latest advances in the field Written for students of life sciences, physics, astronomy and related disciplines, the updated edition of Astrobiology is an essential introductory text that includes recent advances to this dynamic field.

Оглавление

Charles S. Cockell. Astrobiology

Table of Contents

List of Tables

List of Illustrations

Guide

Pages

Astrobiology. Understanding Life in the Universe

Acknowledgments

About the Companion Website

1 Astrobiology. Learning Outcomes

1.1 Introductory Remarks

1.2 The Major Questions of Astrobiology and the Content of the Textbook

1.3 Some Other Features of the Textbook

1.4 A Brief History of Astrobiology

1.5 Conclusions

Bibliography. Books

Papers

2 What Is Life? Learning Outcomes

2.1 The Concept of “Life”

2.2 What Is Life? The Historical Perspective

2.3 Spontaneous Generation

Focus: Astrobiologists: Mary Beth Wilhelm

2.4 More Modern Concepts

2.5 Schrödinger and Life

2.6 Life as a Dissipative Process

2.7 Life: Just a Human Definition?

2.8 Does It Matter Anyway?

2.9 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

3 Matter and Life. Learning Outcomes

3.1 Matter and Life

3.2 Life Is Made of “Ordinary” Matter

3.3 The Atomic Nucleus

3.3.1 Isotopes

3.4 Electrons, Atoms, and Ions

3.5 Types of Bonding in Matter

3.6 Ionic Bonding

3.6.1 Ionic Bonds and Life

3.7 Covalent Bonding

3.7.1 Covalent Bonds and Life

3.8 Metallic Bonding

Discussion Point: Why Doesn't Terrestrial Biochemistry Use Metal Structures?

3.9 Van der Waals Interactions

3.9.1 Dipole–Dipole (Keesom) Forces

3.9.2 Dipole-Induced Dipole (Debye; Pronounced Deh-beye) Forces

3.9.3 Dispersion Forces

3.9.4 Van der Waals Interactions and Life

Focus: Astrobiologists: Andreas Elsaesser

3.10 Hydrogen Bonding

3.10.1 Hydrogen Bonds and Life

3.11 An Astrobiological Perspective

Discussion Point: Is the Structure of Life Universal?

3.12 The Equation of State Describes the Relationship Between Different Types of Matter

3.12.1 Phase Diagrams

3.12.2 Phase Diagrams and Mars

3.12.3 Phase Diagrams and Life

3.13 Other States of Matter

3.13.1 Plasma

3.13.2 Degenerate Matter

Discussion Point: Can Life be Made from Different States of Matter to Terrestrial Life?

3.14 The Interaction Between Matter and Light

3.14.1 The Special Case of the Hydrogen Atom

3.14.2 Uses for Astrobiology

3.15 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

4 The Molecular Structure of Life. Learning Outcomes

4.1 Building Life

4.2 The Essential Elements: CHNOPS

Discussion Point: Are the Minimal CHNOPS Elements a Universal Requirement for Life?

4.3 Carbon Is Versatile

4.4 The Chains of Life

Discussion Point: Searching for Life Without Knowing Anything About it

4.5 Proteins

Discussion Point: Why Does Life Use the 20 Particular Amino Acids it Has?

4.6 Chirality

Discussion Point: Life with a Different Chirality

4.7 Carbohydrates (Sugars)

Focus: Astrobiologists: Scott Perl

4.8 Lipids

4.9 The Nucleic Acids

4.9.1 Ribonucleic Acid

Discussion Point: How Different Could the Genetic Code Be?

4.10 The Solvent of Life

4.10.1 Water as a Solvent

4.11 Alternative Chemistries

4.11.1 Alternative Core Elements

4.11.2 Alternative Solvents

4.12 The Structure of Life and Habitability

4.13 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

5 The Cellular Structure of Life. Learning Outcomes

5.1 From Molecules to Cells

5.2 Types of Cells

Discussion Point: Must Life Be Cellular?

5.3 Shapes of Cells

5.4 The Structure of Cells

5.5 The Structure of Cellular Membranes

5.5.1 Gram-Negative and -Positive Prokaryotic Membranes

Discussion Point: Are Prokaryotes “Simple”?

5.5.2 Archaeal Membranes

5.6 The Information Storage System of Life

5.6.1 Transcription – DNA to RNA

5.6.2 Translation – RNA to Protein

Discussion Point: Why Is There Degeneracy in the Genetic Code?

5.6.3 A Remarkable Code

5.6.4 The Evolution of the Codons

Discussion Point: The Universality of the Genetic Structure and Machinery

Focus: Astrobiologists: Nicol Caplin

5.6.5 DNA Replication

5.6.6 Plasmids

Discussion Point: What Is the Minimum Size of a Cell?

5.6.7 eDNA

5.7 Eukaryotic Cells

5.7.1 Endosymbiosis

5.8 The Reproduction of Cells

5.9 Why Did Sexual Reproduction Evolve?

5.10 The Growth of Populations of Cells

5.11 Moving and Communicating

5.11.1 Movement in Prokaryotes

Discussion Point: Rotating Structures in Nature – Why Don't Animals Have Wheels?

5.11.2 Communication in Prokaryotes

5.11.3 “Multicellularity” in Prokaryotes and Single-Celled Eukaryotes

Discussion Point: When Did Multicellularity Evolve?

5.12 Viruses

Discussion Point: Astrobiology and Viruses

5.13 Prions

5.14 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

6 Energy for Life. Learning Outcomes

6.1 Energy and Astrobiology

6.2 Life and Energy

6.3 The Central Role of Adenosine Triphosphate

6.4 Chemiosmosis and Energy Acquisition

Discussion Point: Energy from Something Other Than Electrons?

6.5 What Types of Electron Donors and Acceptors Can Be Used?

6.6 Aerobic Respiration

6.7 Anaerobic Respiration

6.8 Fermentation

6.9 Chemoautotrophs: Changing the Electron Donor

Focus: Astrobiologists: Haley Sapers

6.9.1 Chemoautotrophy and Fixing CO2

6.9.2 Methanogens and Insights into the Evolution of Metabolism

6.9.3 How Universal Is Metabolism? The TCA Cycle Revisited

Discussion Point: Are Metabolic Pathways Universal?

6.9.4 Methanotrophs

Discussion Point: Geochemical Disequilibria, Planets, and Life

6.9.5 Sulfur Cycling

6.9.6 Iron Oxidizers

6.9.7 Nitrogen Cycling and the Chemoautotrophs

6.10 Energy from Light: Photosynthesis

6.11 Oxygenic Photosynthesis

Discussion Point: Is Oxygenic Photosynthesis Inevitable?

6.12 Anoxygenic Photosynthesis

6.13 Rhodopsins and Photosynthesis

6.14 Evolution of Photosynthesis

6.15 Global Biogeochemical Cycles

Discussion Point: Life Independent of Photosynthesis

6.16 Microbial Mats – Energy-Driven Zonation in Life

6.17 The Thermodynamics of Energy Acquisition and Life

6.17.1 Gibbs Free Energy: The Energy in Reactants and Products

6.17.2 Gibbs Free Energy: The Concentration of Compounds

6.17.3 Gibbs Free Energy: Using Redox Reactions

6.17.4 Gibbs Free Energy: An Example

6.18 Energy and Life in Extremes

6.19 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

7 The Limits of Life. Learning Outcomes

7.1 The Limits of Life

7.2 The Importance of the Limits of Life for Astrobiology

7.3 The Most Extreme Conditions are Dominated by Microbes

7.4 Life at High Temperatures

Focus: Astrobiologists: Alexandra Pontefract

7.4.1 Uses for Thermostable Molecules

7.5 Life at Low Temperatures

Facilities Focus: Underground Laboratories

7.6 Salt-Loving Organisms

7.6.1 Salt-In Strategy

7.6.2 Salt-Out Strategy

7.6.3 Low Water Activity

Discussion Point: Salts in the Universe

7.7 pH Extremes

7.8 Life Under High Pressure

Focus: Astrobiologists: David Smith

7.9 Tolerance to High Radiation

7.10 Life in Toxic Brews

7.11 Rocks as a Habitat

Facilities Focus: Labs in Other Extreme Environments

7.12 Polyextremophiles – Dealing with Multiple Extremes

Discussion Point: The Limits of Life – Are They Universal?

7.13 Life Underground

7.14 Dormancy in Extreme Conditions

7.15 Eukaryotic Extremophiles

7.16 Are There Other Biospheres with Different Limits?

7.17 The Limits of Life: Habitability Revisited

7.18 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

8 The Tree of Life. Learning Outcomes

8.1 A Vast Quantity of Life

Discussion Point: How Much Life Is There on Earth?

8.2 Evolution and a “Tree of Life”

8.3 Classifying Organisms

8.4 The Tree of Life and Some Definitions

Discussion Point: Must All Life Be Assembled into Some Type of Tree?

8.5 Problems with Classification: Homology and Analogy

Discussion Point: Vestigiality, Chance, or Function?

8.6 Building a Phylogenetic Tree Using Genetic Material

Focus: Astrobiologists: Sanjoy Som

8.7 Types of Phylogenetic Trees

8.8 A Modern View of the Tree of Life

8.9 Using Phylogenetic Trees to Test Hypotheses

8.10 Complications in Building Trees

8.10.1 Endosymbiosis

8.10.2 Horizontal Gene Transfer

Discussion Point: Is the Tree of Life a Mirage?

8.11 Origin of Eukaryotes

8.12 The Last Universal Common Ancestor

Discussion Point: Can We Ever Know the LUCA?

8.13 Multiple Origins of Life?

Discussion Point: Shadow Biospheres?

8.14 Alien Life

8.15 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

9 The Universe, the Solar System, and the Elements of Life. Learning Outcomes

9.1 Our Cosmic Situation

9.2 In the Beginning: The Formation of the Universe

9.3 Stellar Evolution: Low-Mass Stars

9.4 Stellar Evolution: High-Mass Stars

9.5 The Elements of Life

9.6 The Hertzsprung–Russell Diagram

Facilities Focus: Space Telescopes and Observatories

Focus: Astrobiologists: Jason Wright

9.7 The Sun Is a Blackbody

9.8 The Formation of Planets

9.9 Types of Objects in Our Solar System

9.10 Meteorites and Their Classification

Discussion Point: Old Meteorites and Their Preservation

9.11 Laws Governing the Motion of Planetary Bodies

9.12 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

10 Astrochemistry: Carbon in Space. Learning Outcomes

10.1 Astrochemistry: Carbon Molecules in Space

10.2 Observing Organics

10.3 In the Beginning

10.4 Different Environments for Chemistry

10.4.1 Diffuse Interstellar Clouds

10.4.2 Molecular Clouds

10.4.3 Protoplanetary Discs

10.4.4 Carbon-Rich Stars

10.4.5 Shock Waves from Supernova Explosions

10.5 How Do Chemical Reactions Occur?

10.6 Forming Carbon Compounds

10.7 Formation of Water

Discussion Point: Water as a Solvent for Life

10.8 Interstellar Grains

10.9 Polycyclic Aromatic Hydrocarbons

10.10 Even More Carbon Diversity

10.10.1 Prebiotic Compounds

10.11 Comets and Organic Molecules

10.12 The Origin of Chirality

10.13 Laboratory Experiments

Discussion Point: Replicating Interstellar Conditions in the Laboratory

10.14 Observing Organic Molecules

10.15 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

11 Early Earth: The First Billion Years. Learning Outcomes

11.1 The First Billion Years of Earth

11.2 Earth Forms and Differentiates

11.3 The Formation of the Moon

11.4 The Early Oceans

General Focus: Astrobiologists: Ania Losiak

11.5 The Early Crust

11.6 The Early Atmosphere

11.7 The Temperature of Early Earth

11.8 The Late Heavy Bombardment

Discussion Point: Age Dating Using Craters

11.9 Implications of the Early Environment for Life

Discussion Point: Radiation – Good for Life?

11.10 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

12 The Origin of Life. Learning Outcomes

12.1 The Origin of Life

12.2 The Synthesis of Organic Compounds on Earth

Discussion Point: Is the Origin of Life Inevitable?

12.2.1 Possible Reaction Pathways

12.3 Delivery from the Extraterrestrial Environment

Focus: Astrobiologists: Sarah Stewart Johnson

12.4 The RNA World

Discussion Point: What Came First?

12.5 Early Cells

12.6 Where Did the Origin of Life Occur?

12.6.1 Deep Sea Hydrothermal Vents

12.6.2 Land-Based Volcanic Environments

12.6.3 Impact Craters

12.6.4 On the Beach

12.6.5 Atmospheric Bubbles

12.6.6 The Deep Subsurface

12.6.7 Mineral Surfaces

12.7 A Cold Origin of Life?

12.8 The Whole Earth as a Reactor?

Discussion Point: No Specific Location for the Emergence of the First Cell?

12.9 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

13 Early Life on Earth. Learning Outcomes

13.1 Early Life on Earth

13.2 Early Life – Metabolisms and Possibilities

13.3 Isotopic Fractionation

General Focus: Astrobiologists: Sean McMahon

13.3.1 Carbon Isotopes

13.4 Measuring the Isotope Fractionation: The Delta Notation

13.5 Sulfur Isotope Fractionation

13.6 Using Isotopes to Look for Ancient Life

13.7 Morphological Evidence for Life

13.7.1 How Are Microorganisms Fossilized?

13.7.2 Evidence for Ancient Fossil Microbial Life

General Focus: Astrobiologists: Nicola McLoughlin

13.7.3 Stromatolites

Discussion Point: Stromatolite Structures Without Biology?

Discussion Point: MISS On Mars?

13.8 Biomarkers

13.9 Contamination Is a Problem

13.10 Instruments Used to Look for Life

Facilities Focus: Nano Secondary Ion Mass Spectrometry (Nano-SIMS)

13.11 A Brief Summary

13.12 The Search for Extraterrestrial Life

13.13 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

14 The Geology of a Habitable World. Learning Outcomes

14.1 The Geological History of Earth: A Habitable World

14.2 Minerals and Glasses

14.3 Types of Rocks

14.3.1 Igneous Rocks

General Facilities Focus: Studying Rocks and Planetary Materials

14.3.2 Sedimentary Rocks

14.3.3 Metamorphic Rocks

14.4 The Rock Cycle

Discussion Point: Anthropic Rocks

14.5 The Composition of Earth. 14.5.1 The Core and Earth's Magnetic Field

Discussion Point: Is a Magnetic Field Required for Life?

14.5.2 The Mantle

14.5.3 Earth's Crust and Upper Mantle

14.6 Plate Tectonics

Discussion Point: Plate Tectonics and Planetary Habitability

Discussion Point: Plate Tectonics and the Origin of Life

14.7 Dating the Age of the Earth (and Other Planetary Bodies)

General Focus: Astrobiologists: Feng Tian

14.8 Age-Dating Rocks

14.8.1 Absolute Age Dating of Rocks

14.8.1.1 The Isochron Method of Age Dating

14.8.1.2 Extinct Radioisotopes

14.8.2 Relative Dating of Rocks

14.8.2.1 The Principle of Superposition

14.8.2.2 The Principle of Original Horizontality

14.8.2.3 The Principle of Crosscutting Relationships

14.8.2.4 The Principle of Faunal Succession

14.8.3 Unconformities

14.8.3.1 Non-conformity

14.8.3.2 Angular Unconformity

14.8.3.3 Disconformity

14.9 Geological Timescales

14.10 The Major Classifications of Geological Time

14.11 Some Geological Times and Biological Changes

14.11.1 The Precambrian

14.11.2 The Phanerozoic: The Rise of Animals and Complexity

14.11.2.1 The Paleozoic Era

14.11.2.2 The Mesozoic Era

14.11.2.3 The Cenozoic Era

Discussion Point: The Anthropocene

14.12 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

15 The Co-evolution of Life and a Planet: The Rise of Oxygen. Learning Outcomes

15.1 Dramatic Changes on Earth

15.2 Measuring Oxygen Through Time

15.2.1 Minerals that Form at Low Oxygen Concentrations

15.2.2 Changes in the Oxidation State of Elements

15.2.3 Banded Iron Formations and Their Isotopes

15.2.4 Sulfur Isotope Fractionation

15.3 It Was Not a Simple Rise

15.4 Summarizing the Evidence for the GOE

General Focus: Astrobiologists: Jacob Haqq-Misra

15.5 The Source of Oxygen

15.6 Sinks for Oxygen

15.7 Why Did Atmospheric Oxygen Concentrations Rise?

15.8 Snowball Earth Episodes

Discussion Point: Life on a Present-Day Snowball Earth

15.9 Other Biological Consequences of the Rise of Oxygen

15.10 Oxygen and the Rise of Animals

Discussion Point: The Rise of Oxygen on Other Planets

15.11 Oxygen and the Rise of Intelligence

15.12 Periods of High Oxygen

15.13 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

16 Mass Extinctions. Learning Outcomes

16.1 Extinctions

16.2 What Is Extinction?

16.3 Five Major Mass Extinctions

16.4 Other Extinctions in Earth History

16.5 Causes of Mass Extinction

16.5.1 Volcanic Causes

16.5.2 Sea-Level Changes

16.5.3 Asteroid and Comet Impacts

16.5.4 Biological Changes

16.5.5 Supernova Explosions and Gamma Ray Bursts

16.5.6 Multiple Effects

16.6 The End-Cretaceous Extinction

16.6.1 Evidence for Asteroid Impact

Discussion Point: What if... The K–Pg Extinction Didn't Happen?

16.6.2 Other Mechanisms for Extinction at the K–Pg Boundary

16.7 The Other Four Big Extinctions of the Phanerozoic. 16.7.1 End-Ordovician Mass Extinction

16.7.2 Late Devonian Mass Extinction

16.7.3 The Largest of all Mass Extinctions: The End-Permian Extinction

16.7.4 End-Triassic Mass Extinction

16.8 Do Microorganisms Go Extinct?

16.9 Recovery from Extinction

16.10 Can We Avoid Extinction?

16.11 The Sixth Mass Extinction?

Discussion Point: The Sixth Extinction

16.12 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

17 The Habitability of Planetary Bodies. Learning Outcomes

17.1 What Is “Habitability”?

Discussion Point: Is Habitability a Binary Property?

17.2 The Habitable Zone

17.2.1 Effects of Star Types on Habitability

17.2.2 Continuously Habitable Zone

17.2.3 Pre- and Post-Main Sequence Stars

17.3 Maintaining Temperature Conditions on a Planet Suitable for Water and Life

17.3.1 Effective Temperature and the Greenhouse Effect

17.3.1.1 Calculating the Effective Temperature of a Planet

17.3.1.2 The Greenhouse Effect

Discussion Point: Atmospheric Habitability and Airborne Biospheres

17.3.2 The Carbonate–Silicate Cycle

General Focus: Astrobiologists: Shawn Domagal-Goldman

17.3.3 Habitable Conditions Underground and on Lone Planets?

17.4 Plate Tectonics and Habitability

Discussion Point: The Factors Required for Habitability

17.5 Does the Moon Play a Role in Habitability?

General Focus: Astrobiologists: Robin Wordsworth

17.6 Other Planetary Factors that Influence Habitability

17.7 Surface Liquid Water, Habitability, and Intelligence

17.8 Habitable Environments Need Not Always Contain Life

17.9 Worlds More Habitable than Earth?

17.10 The Anthropic Principle and Habitability

17.11 The Fate of Earth

17.12 The Galactic Habitable Zone

17.13 The Right Galaxy?

17.14 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

18 The Astrobiology of Mars. Learning Outcomes

18.1 Mars and Astrobiology

18.2 Martian Geological History: A Very Brief Summary

18.3 The Environmental Deterioration of Mars

18.4 Missions to Mars

18.5 Mars and Life

18.5.1 Water and Mars. 18.5.1.1 The Evidence for Water on Mars

Discussion Point: Mars – A Better Place to Look for Early Life than Earth?

18.5.1.2 Liquid Water on Mars Today

18.5.2 Basic Elements for Life on Mars

18.5.2.1 Carbon

18.5.2.2 Hydrogen

18.5.2.3 Nitrogen

18.5.2.4 Oxygen

18.5.2.5 Phosphorus

18.5.2.6 Sulfur

18.5.3 Trace Elements for Life on Mars

18.5.4 Energy and Redox Couples for Life on Mars

General Focus: Astrobiologists: Ralf Moeller

18.5.5 Physical Limits to Life: Radiation

18.5.6 Physical Limits to Life: pH

18.5.7 Physical Limits to Life: Salts

18.5.8 Habitat Space for Microbes on Mars

18.6 Trajectories of Martian Habitability

18.6.1 Trajectories for an Uninhabited Mars

General Facilities Focus: Mars Simulation Chambers

18.6.2 Trajectories for an Inhabited Mars

18.7 The Viking Program and the Search for Life

18.7.1 GC–MS Analysis

18.7.2 Gas-Exchange Experiment

18.7.3 Labeled Release Experiment

18.7.4 Pyrolytic Release Experiment

18.7.5 Viking: A Lesson in Science

18.8 Searching for Life by Investigating Gases

18.9 Martian Meteorites

18.10 Mars Analog Environments

Discussion Point: Astrobiology, Extraterrestrial Life, and Science

18.11 Panspermia: The Transfer of Life between Planets?

18.11.1 Ejection from a Planet

18.11.2 Interplanetary Transfer Phase

18.11.3 Arriving at the Destination Planet

18.12 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

19 Ocean Worlds and Icy Moons. Learning Outcomes

19.1 The Astrobiology of Moons

19.2 The Moons of Jupiter: Europa

Discussion Point: Plate Tectonics on Europa

19.2.1 A Subsurface Ocean

19.2.1.1 Habitable Conditions in Europa's Ocean?

19.3 The Moons of Jupiter: Ganymede and Callisto

19.4 The Moons of Jupiter: Io

19.5 The Moons of Saturn: Enceladus

general Focus: Astrobiologists: Nozair Khawaja

19.5.1 The Plumes of Enceladus

Discussion Point: The Exploration of Ocean Worlds

19.6 The Moons of Saturn: Titan

19.7 Other Icy Worlds

19.7.1 Ceres

19.7.2 Neptune's Moon Triton

19.7.3 Pluto

19.8 Planetary Protection

Discussion Point: Is Planetary Protection Necessary?

19.9 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

20 Exoplanets and the Search for Life. Learning Outcomes

20.1 Exoplanets and Life

20.2 Detecting Exoplanets. 20.2.1 The Challenge of Detection

20.2.2 Transit Method

20.2.3 Doppler Shift/Radial Velocity Method

20.2.4 Astrometry

General Focus: Astrobiologists: Sarah Rugheimer

20.2.5 Variations in Other Attributes of Stars

20.2.6 Orbital Brightness Changes

20.2.7 Gravitational Lensing

20.2.8 Direct Detection

20.2.9 Using Direct Detection to Study Protoplanetary Discs

20.3 Exoplanet Properties. 20.3.1 General Properties

20.3.2 Hot Jupiters and Neptunes

20.3.3 Super-Earths and Ocean Worlds

20.3.3.1 Plate Tectonics and Habitability on Super-Earths and Ocean Worlds

20.3.4 Rocky Planets in the Habitable Zone

Discussion Point: Rare Earth?

20.3.5 Planets in Binary and Multiple Star Systems

20.3.6 Strange Worlds

20.3.7 Habitable Exomoons?

General Focus: Astrobiologists: Eddie Schwieterman

20.4 Detecting Life. 20.4.1 Biosignature Gases

20.4.2 Oxygen as a Biosignature Gas and Atmospheric Disequilibria with Oxygen

Discussion Point: How Universal Are Biosignature Gases?

20.4.3 False Positives and False Negatives

20.4.4 Anti-Biosignature Gases

20.4.5 Other Biosignature Gases

20.4.6 Technosignature Gases

20.5 Surface Biosignatures

Discussion Point: What Happens after a Biosignature Detection?

20.6 How Likely Are These Signatures?

20.7 Other Ways to Find Life

20.8 Missions to Detect Biosignatures

20.9 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

21 The Search for Extraterrestrial Intelligence. Learning Outcomes

21.1 The Search for Extraterrestrial Intelligence (SETI)

21.2 Methods in the Search for Extraterrestrial Intelligence

21.2.1 Searching in the Radio Region of the Electromagnetic Spectrum

21.2.2 Searching in the Optical Region of the Electromagnetic Spectrum

21.2.3 The Significance of Exoplanet Research

21.2.4 The Search for Extraterrestrial Artifacts (SETA)

21.3 Communication with Extraterrestrial Intelligence (CETI)

21.4 The Drake Equation

Discussion Point: The Drake Equation

21.5 The Fermi Paradox

21.5.1 Civilizations Are Too Far Apart in Space

21.5.2 No Other, or Very Few, Civilizations Have Arisen

Discussion Point: Is the Fermi Paradox a Scientific Hypothesis?

21.5.3 Intelligent Life Destroys Itself

21.5.4 Civilizations Are Periodically Destroyed by Natural Events

21.5.5 It Is the Nature of Intelligent Life to Destroy Other Civilizations

21.5.6 They Exist, But We See No Evidence of Them

21.5.7 They Are in the Local Area, But Observing Us Rather than Attempting to Make Contact

21.5.8 They Are Too Busy Online

21.5.9 They Are Here

21.5.10 The Evidence Is Being Suppressed

21.6 Classifying Civilizations

21.7 Policy Implications

21.8 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

22 Our Civilization. Learning Outcomes

22.1 Astrobiology and Human Civilization

22.2 The Emergence of Human Society

22.3 Threats to a Civilization

22.3.1 Supervolcanoes

22.3.2 Asteroid and Comet Impacts

22.3.3 Disease

22.4 Climate Change and the Challenge to Civilization

22.5 The Human Future Beyond Earth

22.5.1 The Rocket Equation

22.6 Settling the Solar System. 22.6.1 Where First?

Discussion Point: What Does It Take to Live Self-Sufficiently on the Moon and Mars?

22.6.2 Life Support Systems

22.6.2.1 Compartment 1: The Liquefying Compartment

22.6.2.2 Compartment 2: The Photoheterotrophic Compartment

22.6.2.3 Compartment 3: The Nitrifying Compartment

22.6.2.4 Compartment 4: The Photoautotrophic Compartment

General Focus: Astrobiologists: Octavio A. Chon Torres

22.6.3 Human Exploration of Mars

Discussion Point: Terraforming

22.6.4 The Asteroids

22.6.5 The Outer Solar System

22.6.6 Large Human Settlements in Space

22.7 Avoiding Extinction or Collapse: A Multiplanet Species

22.8 Environmentalism and Space Exploration as a Single Goal?

22.9 Sociology: The Overview Effect

22.10 Will We Become Interstellar?

22.11 Conclusions

Questions for Review and Reflection

Bibliography. Books

Papers

Appendix. A.1 The Astrobiology Periodic Table

A.2 Units and Scales. A.2.1 Standard International Base Units

A.2.2 Basic Physical Constants

A.3 Temperature Scale Conversion

A.4 Composition of the Sun

A.5 Some of the Major Star Types, Temperatures, and Colors

A.6 Three- and One-Letter Designations of Amino Acids

A.7 Codon Table for the Genetic Code Associated with mRNA (also shown in Chapter 5; Figure 5.12)

A.8 Planetary Data

A.9 Geological Time Scale

Glossary

Index

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Second Edition

CHARLES S. COCKELL

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In 1960, Joshua Lederberg (1925–2008; Figure 1.16), a pioneer in bacteriology and molecular biology who won the Nobel Prize for his work on bacterial genetics, used the term exobiology to describe the search for life beyond Earth. This term later became associated specifically with the search for life beyond Earth, which now constitutes just one word in the wider term: astrobiology. Other terms that appear in the popular and scientific literature have included cosmobiology, xenobiology, and bioastronomy, the latter used mainly by astronomers.

Figure 1.16 Nobel Laureate Joshua Lederberg. He was at the forefront of the United States' efforts in exobiology in the twentieth century, at his laboratory in the University of Wisconsin, October 1958.

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