Hadrosaurs
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Оглавление
David A. Eberth. Hadrosaurs
HADROSAURS
C. Contents
C. Contributors
R. Reviewers
P. Preface
A. Acknowledgments
1. A History of the Study of Ornithopods: Where Have We Been? Where Are We Now? and Where Are We Going?
ABSTRACT
INTRODUCTION
MATERIALS AND METHODS
WHERE HAVE WE BEEN?
WHERE ARE WE NOW?
WHERE ARE WE GOING?
ACKNOWLEDGMENTS
LITERATURE CITED
2. Iguanodonts from the Wealden of England: Do They Contribute to the Discussion Concerning Hadrosaur Origins?
ABSTRACT
INTRODUCTION
A REVIEW OF WEALDEN IGUANODONTIANS
SYSTEMATIC PALEONTOLOGY
Taxonomic Discussion
Description
Reconstruction of Barilium
Taxonomic Discussion
Description
Reconstruction of Hypselospinus
Taxonomic Discussion
Description
Reconstruction of Mantellisaurus
Taxonomic Discussion
Description
Reconstruction of Iguanodon bernissartensis
SYSTEMATIC ANALYSIS
Data and Methodological Framework
Results
CHARACTER DISTRIBUTION AND PHYLOGENETIC IMPLICATIONS
Basal Ornithopod Taxa
Clypeodonta clade nov
Character Acquisition at This Node
Condensed Diagnosis
Dryomorpha
Character Acquisition at This Node
Condensed Diagnosis
Ankylopollexia
Character Acquisition at This Node
Condensed Diagnosis
Neoiguanodontia (“New Iguanodonts”)
Character Acquisition at This Node
Condensed Diagnosis
Hadrosauromorpha clade nov
Character Acquisition between Probactrosaurus and the Succeeding Node
Condensed Diagnosis
Euhadrosauria
Abbreviated Diagnosis
A DISCUSSION: THE ORIGIN OF HADROSAURS
Topological Variation
An Evolutionary Narrative of Hadrosaur Origins
ACKNOWLEDGMENTS AND DEDICATION
LITERATURE CITED
3. Osteology of the Basal Hadrosauroid Equijubus normani (Dinosauria, Ornithopoda) from the Early Cretaceous of China
ABSTRACT
INTRODUCTION
SYSTEMATIC PALEONTOLOGY
DESCRIPTION. Cranium and Mandible
Axial Skeleton
Appendicular Skeleton
DISCUSSION. Ontogenetic Age of IVPP V 12534
Changes Made to Previous Diagnoses of Equijubus
Phylogenetic Relationships of Equijubus
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
4. Gongpoquansaurus mazongshanensis (Lü, 1997) comb. nov. (Ornithischia: Hadrosauroidea) from the Early Cretaceous of Gansu Province, Northwestern China
ABSTRACT
INTRODUCTION
SYSTEMATIC PALEONTOLOGY
DISCUSSION
ACKNOWLEDGMENTS
LITERATURE CITED
5. Postcranial Anatomy of a Basal Hadrosauroid (Dinosauria: Ornithopoda) from the Cretaceous (Cenomanian) Woodbine Formation of North Texas
ABSTRACT
INTRODUCTION
GEOLOGIC SETTING. Woodbine Formation and Members
Stratigraphy and Paleoenvironments of the AAS
Taphonomy
SYSTEMATIC PALEONTOLOGY
DESCRIPTION
DISCUSSION. Taxonomic Affinity
Comparison with Woodbine Ornithopods
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
6. A Re-evaluation of Purported Hadrosaurid Dinosaur Specimens from the “Middle” Cretaceous of England
ABSTRACT
INTRODUCTION
DESCRIPTIONS “Trachodon cantabrigiensis” Lydekker, 1888a
‘Iguanodon’ hillii Newton, 1892
DISCUSSION. Morphological Comparisons
Character Analysis and Phylogenetic Positions of “Trachodon cantabrigiensis” and ‘Iguanodon’ hillii
Morphometric Analysis
Validity and Affinities of English “Hadrosaurids”
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
7. A New Hadrosauroid (Plesiohadros djadokhtaensis) from the Late Cretaceous Djadokhtan Fauna of Southern Mongolia
ABSTRACT
INTRODUCTION
GEOLOGICAL SETTING
Sedimentological Environments of the Djadokhta Formation
Age of the Djadokhta Formation
SYSTEMATIC PALEONTOLOGY
DESCRIPTION
Lower Jaw
Dentition
Postcranium
PHYLOGENETIC ANALYSIS. Methods
Results
DISCUSSION
ACKNOWLEDGMENTS
LITERATURE CITED
8. Hadrosauroid Material from the Santonian Milk River Formation of Southern Alberta, Canada
ABSTRACT
INTRODUCTION
GEOLOGICAL SETTING
SYSTEMATIC PALEONTOLOGY
DESCRIPTION. Cranial Bones
Postcranial Bones
PHYLOGENETIC ANALYSIS. Methods
Results
DISCUSSION. Affinities and Implications of the Milk River Hadrosauroid Material
Biogeographic Implications
ACKNOWLEDGMENTS
LITERATURE CITED
9. New Hadrosaurid (Dinosauria, Ornithopoda) Specimens from the Lower–Middle Campanian Wahweap Formation of Southern Utah
ABSTRACT
INTRODUCTION
SYSTEMATIC PALEONTOLOGY
Description
Description
Femoral Histology and Ontogenetic Status of UMNH VP 20187
Taxonomic Identification
Phylogenetic Analyses
Taphonomy of the WBH Locality
DISCUSSION. Significance of Adelolophus
WBH Locality Behavioral Implications
Wahweap Hadrosaurid Diversity
ACKNOWLEDGMENTS
LITERATURE CITED
10. New Saurolophine Material from the Upper Campanian–Lower Maastrichtian Wapiti Formation, West-Central Alberta
ABSTRACT
INTRODUCTION
LOCALITY AND GEOLOGICAL SETTING
SYSTEMATIC PALEONTOLOGY
Description
HISTOLOGY. Methods
Results
PHYLOGENETIC ANALYSIS. Methods
Results
DISCUSSION. Ontogenetic Status of TMP 1989.092.0001
Evolutionary and Biogeographic Implications
ACKNOWLEDGMENTS
LITERATURE CITED
11. Variation in the Skull Roof of the Hadrosaur Gryposaurus Illustrated by a New Specimen from the Kaiparowits Formation (late Campanian) of Southern Utah
ABSTRACT
INTRODUCTION
SYSTEMATIC PALEONTOLOGY
Description
STATISTICAL ANALYSIS. Methods
Results
DISCUSSION
ACKNOWLEDGMENTS
LITERATURE CITED
12. A Skull of Prosaurolophus maximus from Southeastern Alberta and the Spatiotemporal Distribution of Faunal Zones in the Dinosaur Park Formation
ABSTRACT
INTRODUCTION
SYSTEMATIC PALEONTOLOGY
DESCRIPTION
DISCUSSION
ACKNOWLEDGMENTS
LITERATURE CITED
13. Postcranial Anatomy of Edmontosaurus regalis (Hadrosauridae) from the Horseshoe Canyon Formation, Alberta, Canada
ABSTRACT
INTRODUCTION
SYSTEMATIC PALEONTOLOGY
DESCRIPTION. Axial Skeleton
Pectoral Girdle
Forelimb
Pelvic Girdle
Hindlimb
DISCUSSION. Pelvic Morphology
Vertebral Variation
Limb Proportions and Body Mass Estimates of Edmontosaurus regalis
ACKNOWLEDGMENTS
LITERATURE CITED
14. Cranial Morphology and Variation in Hypacrosaurus stebingeri (Ornithischia: Hadrosauridae)
ABSTRACT
INTRODUCTION
MATERIALS AND METHODS
SYSTEMATIC PALEONTOLOGY
HYPACROSAURUS STEBINGERI Horner and Currie, 1994
DESCRIPTION
Maxillary Complex
Neurocranial Complex
MORPHOMETRIC ANALYSIS
DISCUSSION
ACKNOWLEDGMENTS
LITERATURE CITED
15. An Overview of the Latest Cretaceous Hadrosauroid Record in Europe
ABSTRACT
INTRODUCTION
TERMINOLOGY AND METHODS
THE LATEST CRETACEOUS HADROSAUROID RECORD OF EUROPE
Ukraine
Bulgaria
Romania
Slovenia
Italy
Germany
Belgium and the Netherlands
France
Spain
DISCUSSION. Body Size of European Hadrosauroids and Insular Dwarfism
Hadrosauroid Diversity in the European Archipelago During the Latest Cretaceous
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
16. The Hadrosauroid Record in the Maastrichtian of the Eastern Tremp Syncline (Northern Spain)
ABSTRACT
INTRODUCTION
METHODS AND TERMINOLOGY
EASTERN TREMP SYNCLINE: GEOLOGICAL SETTING
HADROSAUROID SITES AND REMAINS IN THE EASTERN TREMP SYNCLINE
Hadrosauroid Bone-Bearing sites
Hadrosauroid Track-Bearing Sites
Dinosaur Eggshell-Bearing Sites
DISCUSSION
Environmental Preference or Time/Event Related Faunal Change?
Why Are There No Hadrosauroid Eggshell Remains?
Are the Hadrosauroids from the Eastern Tremp Syncline Small-Sized and Insular Dwarfs?
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
17. Hadrosaurs from the Far East: Historical Perspective and New Amurosaurus Material from Blagoveschensk (Amur Region, Russia)
ABSTRACT
INTRODUCTION
THE FIRST HADROSAUR DISCOVERIES IN THE AMUR REGION: A HISTORICAL PERSPECTIVE
THE BLAGOVESCHENSK DINOSAUR LOCALITY
NEW AMUROSAURUS MATERIAL FROM BLAGOVESCHENSK AND ITS IMPORTANCE. Humerus AEHM 1/1755: What Was the Maximal Size of Amurosaurus riabinini?
Premaxilla AEHM 1/1038: The Crest of Amurosaurus riabinini
Predentary AEHM 1/1682
Sacra AEHM 1/1715, AEHM 1/1716
PHYLOGENETIC POSITION OF AMUROSAURUS RIABININI
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
18. South American Hadrosaurs: Considerations on Their Diversity
ABSTRACT
INTRODUCTION
SOUTH AMERICAN HADROSAURIDS
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
19. The Hadrosaurian Record from Mexico
ABSTRACT
INTRODUCTION
STRATIGRAPHY AND PALEOENVIRONMENTS
Mojado Formation
Barranca Los Bonetes Area
Rosario Area (La Bocana Roja Formation and El Gallo Formation)
Cabullona Group (Corral de Enmedio, Camas, Packard and Lomas Coloradas formations)
Aguja Formation
Olmos Formation
Difunta Group: Cerro del Pueblo and Cerro Huerta Formations
Mexcala Formation
Arenisca Aguililla Unit
THE HADROSAUROID FOSSIL RECORD OF MEXICO. Cerro Del Cristo Rey Site, Chihuahua State
Barranca Los Bonetes Area, Michoacán State
El Rosario Area and Eréndira Area, Baja California State
Cabullona Group, Sonora State
Aguja Formation, Northeastern Chihuahua State and Northwestern Coahuila State
Ojinaga Area, Northeastern Chihuahua State
Jiménez Area, Southeastern Chihuahua State
Parras Basin: Huerta Formation and Cerro del Pueblo Formation, Coahuila State
Unknown Formation, Coahuila State
Sabinas Basin: Olmos Formation and Escondido Formation (Coahuila)
Mexcala Formation, Puebla State
Arenisca Aguililla Unit, Michoacán State
CONCLUSION
ACKNOWLEDGMENTS
LITERATURE CITED
20. Stratigraphic Distribution of Hadrosaurids in the Upper Cretaceous Fruitland, Kirtland, and Ojo Alamo Formations, San Juan Basin, New Mexico
ABSTRACT
INTRODUCTION
GEOLOGIC AND BIOSTRATIGRAPHIC OVERVIEW
THE SAN JUAN BASIN HADROSAURID-BEARING FORMATIONS. Fruitland Formation
Fruitland–Kirtland Transition
Kirtland Formation (Kirtland Shale)
Ojo Alamo Formation (Ojo Alamo Sandstone)
LOCAL FAUNAS
Hunter Wash Local Fauna
Willow Wash Local Fauna
Alamo Wash Local Fauna
HADROSAURIDS FROM THE SAN JUAN BASIN: AN OVERVIEW
STRATIGRAPHIC DISTRIBUTION OF THE SAN JUAN BASIN HADROSAURIDS
TAXONOMIC DIVERSITY OF THE SAN JUAN BASIN HADROSAURIDS
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
NOTE ADDED IN PROOF
REFERENCE
21. Relocating the Lost Gryposaurus incurvimanus Holotype Quarry, Dinosaur Provincial Park, Alberta, Canada
ABSTRACT
INTRODUCTION
RELOCATION EFFORT
SCIENTIFIC SIGNIFICANCE
CONCLUSIONS
FUTURE PROJECT PERSPECTIVES
ACKNOWLEDGMENTS
LITERATURE CITED
22. Comparative Ontogenies (Appendicular Skeleton) for Three Hadrosaurids and a Basal Iguanodontian: Divergent Developmental Pathways in Hadrosaurinae and Lambeosaurinae
ABSTRACT
INTRODUCTION
MATERIALS AND SPECIMENS
DESCRIPTIVE APPENDICULAR ONTOGENIES. Maiasaura peeblesorum
Hypacrosaurus stebingeri
Brachylophosaurus canadensis
Tenontosaurus tilletti
DISCUSSION
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
23. The Size-Frequency Distribution of Hadrosaurs from the Dinosaur Park Formation of Alberta, Canada
ABSTRACT
INTRODUCTION
METHODS
RESULTS
DISCUSSION
ACKNOWLEDGMENTS
LITERATURE CITED
24. Osteohistology and Occlusal Morphology of Hypacrosaurus stebingeri Teeth throughout Ontogeny with Comments on Wear-Induced Form and Function
ABSTRACT
INTRODUCTION
MATERIALS AND METHODS
RESULTS. Embryonic and Hatchling Teeth
Subadult Teeth
Adult Teeth
DISCUSSION
ACKNOWLEDGMENTS
LITERATURE CITED
25. Three-Dimensional Computational Modeling of Pelvic Locomotor Muscle Moment Arms in Edmontosaurus (Dinosauria, Hadrosauridae) and Comparisons with Other Archosaurs
ABSTRACT
INTRODUCTION
MATERIALS AND METHODS. Myological Reconstruction
Model Construction
Sensitivity Analysis
Comparative Analysis
RESULTS. Edmontosaurus Moment Arms
Sensitivity Analysis
Comparative Moment Arms
DISCUSSION. Muscle Leverage in Edmontosaurus
Functional and Evolutionary Trends and Implications
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
26. Duckbills on the Run: The Cursorial Abilities of Hadrosaurs and Implications for Tyrannosaur-Avoidance Strategies
ABSTRACT
THE MYSTERY OF SUCCESSFUL DUCKBILLS
MATERIALS, METHODS, AND QUANTITATIVE RESULTS
TAIL ANATOMY
DISCUSSION. The Biomechanical Thought Experiment of Racing Dinosaurs
Endurance Running as a Predation Defense Strategy
ACKNOWLEDGMENTS
LITERATURE CITED
27. Duck Soup: The Floating Fates of Hadrosaurs and Ceratopsians at Dinosaur Provincial Park
ABSTRACT
INTRODUCTION
METHODS. Body Model Generation
Assignment of Body Densities
Determination of Flotation Equilibrium
RESULTS
DISCUSSION
CONCLUSION
LITERATURE CITED
28. Hadrosauroid Jaw Mechanics and the Functional Significance of the Predentary Bone
ABSTRACT
INTRODUCTION
MATERIALS AND METHODS
DESCRIPTION
Predentary
Dentary
Splenial
Angular
Surangular, Articular, and Their Association with the Quadrate
Dental Micro- and Mesowear
DISCUSSION. Feeding Mechanism
CONCLUSION
ACKNOWLEDGMENTS
LITERATURE CITED
29. Debris Flow Origin of an Unusual Late Cretaceous Hadrosaur Bonebed in the Two Medicine Formation of Western Montana
ABSTRACT
INTRODUCTION
GEOLOGIC SETTING
METHODS. Fossil Assemblage
Sedimentology and Stratigraphy
RESULTS. Assemblage Data
Sedimentologic Attributes of the Bonebed
DISCUSSION. Bonebed Sedimentology
Skeletal Assemblage Modification
Debris Flows in Alluvial Settings
Origin of the Assemblage
Alternative Hypotheses
CONCLUSION
ACKNOWLEDGMENTS
LITERATURE CITED
30. Occurrence and Taphonomy of the First Documented Hadrosaurid Bonebed from the Dinosaur Park Formation (Belly River Group, Campanian) at Dinosaur Provincial Park, Alberta, Canada
ABSTRACT
INTRODUCTION
METHODS
LOCATION AND GEOLOGY
Interpretation
ASSEMBLAGE DATA
Interpretation
ELEMENT DISTRIBUTIONS AND ORIENTATIONS
ELEMENT MODIFICATIONS
Common Element Modifications
Uncommon Element Modifications
Interpretation
DISCUSSION AND CONCLUSIONS
SUMMARY
ACKNOWLEDGMENTS
LITERATURE CITED
31. Body Size Distribution in a Death Assemblage of a Colossal Hadrosaurid from the Upper Cretaceous of Zhucheng, Shandong Province, China
ABSTRACT
INTRODUCTION
LOCALITY INFORMATION
TAXONOMIC IDENTITY
HADROSAUR MATERIAL AT KUGOU QUARRY
METHODS AND RESULTS
DISCUSSION
SUMMARY
ACKNOWLEDGMENTS
LITERATURE CITED
32. First Hadrosaur Trackway from the Upper Cretaceous (Late Campanian) Oldman Formation, Southeastern Alberta
ABSTRACT
INTRODUCTION
MATERIALS AND METHODS
GEOLOGIC SETTING
THE MILK RIVER NATURAL AREA TRACK SITE
DISCUSSION AND CONCLUSION
ACKNOWLEDGMENTS
LITERATURE CITED
33. Paleopathology in Late Cretaceous Hadrosauridae from Alberta, Canada
ABSTRACT
INTRODUCTION
MATERIALS AND METHODS
PREVIOUS WORK
DESCRIPTION. Skull and Teeth
Vertebrae
Ribs
Shoulder Girdle and Forelimb
Pelvic Girdle and Hindlimb
Ossified Tendons
Eggs and Skin
DISCUSSION
SUMMARY AND CONCLUSIONS
FUTURE DIRECTIONS
ACKNOWLEDGMENTS
LITERATURE CITED
34. A Review of Hadrosaurid Skin Impressions
ABSTRACT
INTRODUCTION
METHODS
DESCRIPTION OF INTEGUMENT IN HADROSAURIDAE. Saurolophinae (sensu Prieto-Márquez, 2010)
Lambeosaurinae (sensu Prieto-Márquez, 2010)
Hadrosauridae indet
DISCUSSION. Characterization of Hadrosaurid Skin Impressions
Comments on Variation and Conservatism in Scale Architecture
CONCLUSIONS
ACKNOWLEDGMENTS
LITERATURE CITED
35. Soft-Tissue Structures of the Nasal Vestibular Region of Saurolophine Hadrosaurids (Dinosauria, Ornithopoda) Revealed in a “Mummified” Specimen of Edmontosaurus annectens
ABSTRACT
INTRODUCTION
DESCRIPTION
DISCUSSION AND CONCLUSION
ACKNOWLEDGMENTS
LITERATURE CITED
36. The Role and Biochemistry of Melanin Pigment in the Exceptional Preservation of Hadrosaur Skin
ABSTRACT
INTRODUCTION
MULTIDISCIPLINARY APPROACHES TO STUDYING EXCEPTIONAL PRESERVATION
SYNCHROTRON RAPID SCANNING X-RAY FLUORESCENCE
RESULTS
DISCUSSION. Pigment of Our Imagination?
Trace-Metal Jacket
CONCLUSION
ACKNOWLEDGMENTS
LITERATURE CITED
A. Afterword
REFERENCES CITED
I. Subject Index
I. Locality Index (by country)
I. Stratigraphy Index (by country)
I. Taxonomic Index
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2. The posterodistal corner of dentary crowns, when viewed lingually, exhibit inrolling of the marginal denticulate edge to form an oblique ledge-like structure (sometimes referred to as a cingulum).
3. Posterior dentary dentition extends medial to the coronoid process, and from which the dentition is separated by a narrow, horizontal ledge that represents a posterior extension of the lateral cheek recess.
.....