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Nerve-endings in the Skin.—"Many of the afferent skin-nerves end in connection with hair-bulbs; the fine hairs over most of the cutaneous surface, projecting from the skin, transmit any movement impressed on them, with increased force, to the nerve-fibres at their fixed ends. Fine branches of axis-cylinders have also been described as penetrating between epidermic cells and ending there without terminal organs. In or immediately beneath the skin several peculiar forms of nerve end-organs have also been described; they are known as (1) Touch-cells; (2) Pacinian corpuscles; (3) Tactile corpuscles; (4) End-bulbs."[23]

Fig. 24.—End-bulbs from the conjunctiva of the human eye, magnified.

These bodies all consist essentially of granules formed of connective tissue, in which or round about which one or more sensory nerve-fibres terminate. They probably magnify impressions just as a grain of sand does in a shoe, or a crumb does in a finger of a glove.

Touch, or the Pressure Sense.—"Through the skin we get several kinds of sensation; touch proper, heat and cold, and pain; and we can with more or less accuracy localize them on the surface of the body. The interior of the mouth possesses also three sensibilities. Through touch proper we recognize pressure or traction exerted on the skin, and the force of the pressure; the softness or hardness, roughness or smoothness, of the body producing it; and the form of this when not too large to be felt all over. When to learn the form of an object we move the hand over it, muscular sensations are combined with proper tactile, and such a combination of the two sensations is frequent; moreover, we rarely touch anything without at the same time getting temperature sensations; therefore pure tactile feelings are rare. From an evolution point of view, touch is probably the first distinctly differentiated sensation, and this primary position it still largely holds in our mental life."[24]

Objects are most important to us when in direct contact. The chief function of our eyes and ears is to enable us to prepare ourselves for contact with approaching bodies, or to ward such contact off. They have accordingly been characterized as organs of anticipatory touch.

"The delicacy of the tactile sense varies on different parts of the skin; it is greatest on the forehead, temples, and back of the forearm, where a weight of 2 milligr. pressing on an area of 9 sq. millim. can be felt.

"In order that the sense of touch may be excited neighboring skin-areas must be differently pressed. When the hand is immersed in a liquid, as mercury, which fits into all its inequalities and presses with practically the same weight on all neighboring immersed areas, the sense of pressure is only felt at a line along the surface, where the immersed and non-immersed parts of the skin meet.

The Localizing Power of the Skin.—"When the eyes are closed and a point of the skin is touched we can with some accuracy indicate the region stimulated; although tactile feelings are in general characters alike, they differ in something besides intensity by which we can distinguish them; some sub-sensation quality not rising definitely into prominence in consciousness must be present, comparable to the upper partials determining the timbre of a tone. The accuracy of the localizing power varies widely in different skin regions and is measured by observing the least distance which must separate two objects (as the blunted points of a pair of compasses) in order that they may be felt as two. The following table illustrates some of the differences observed:

Tongue-tip	1.1 mm.	(.04 inch)
Palm side of last phalanx of finger	2.2 mm.	(.08 inch)
Red part of lips	4.4 mm.	(.16 inch)
Tip of nose	6.6 mm.	(.24 inch)
Back of second phalanx of finger	11.0 mm.	(.44 inch)
Heel	22.0 mm.	(.88 inch)
Back of hand	30.8 mm.	(1.23 inches)
Forearm	39.6 mm.	(1.58 inches)
Sternum	44.0 mm.	(1.76 inches)
Back of neck	52.8 mm.	(2.11 inches)
Middle of back	66.0 mm.	(2.64 inches)

The localizing power is a little more acute across the long axis of a limb than in it; and is better when the pressure is only strong enough to just cause a distinct tactile sensation than when it is more powerful; it is also very readily and rapidly improvable by practice." It seems to be naturally delicate in proportion as the skin which possesses it covers a more movable part of the body.

Fig. 25.

"It might be thought that this localizing power depended directly on nerve-distribution; that each touch-nerve had connection with a special brain-centre at one end (the excitation of which caused a sensation with a characteristic local sign), and at the other end was distributed over a certain skin-area, and that the larger this area the farther apart might two points be and still give rise to only one sensation. If this were so, however, the peripheral tactile areas (each being determined by the anatomical distribution of a nerve-fibre) must have definite unchangeable limits, which experiment shows that they do not possess. Suppose the small areas in Fig. 25 to each represent a peripheral area of nerve-distribution. If any two points in c were touched we should according to the theory get but a single sensation; but if, while the compass-points remained the same distance apart, or were even approximated, one were placed in c and the other on a contiguous area, two fibres would be stimulated and we ought to get two sensations; but such is not the case; on the same skin-region the points must be always the same distance apart, no matter how they be shifted, in order to give rise to two just distinguishable sensations.

"It is probable that the nerve-areas are much smaller than the tactile; and that several unstimulated must intervene between the excited, in order to produce sensations which shall be distinct. If we suppose twelve unexcited nerve-areas must intervene, then, in Fig. 25, a and b will be just on the limits of a single tactile area; and no matter how the points are moved, so long as eleven, or fewer, unexcited areas come between, we would get a single tactile sensation; in this way we can explain the fact that tactile areas have no fixed boundaries in the skin, although the nerve-distribution in any part must be constant. We also see why the back of a knife laid on the surface causes a continuous linear sensation, although it touches many distinct nerve-areas. If we could discriminate the excitations of each of these from that of its immediate neighbors we should get the sensation of a series of points touching us, one for each nerve-region excited; but in the absence of intervening unexcited nerve-areas the sensations are fused together.

The Temperature-sense. Its Terminal Organs.—"By this we mean our faculty of perceiving cold and warmth; and, with the help of these sensations, of perceiving temperature differences in external objects. Its organ is the whole skin, the mucous membrane of mouth and fauces, pharynx and gullet, and the entry of the nares. Direct heating or cooling of a sensory nerve may stimulate it and cause pain, but not a true temperature-sensation; hence we assume the presence of temperature end-organs. [These have not yet been ascertained anatomically. Physiologically, however, the demonstration of special spots in the skin for feeling heat and cold is one of the most interesting discoveries of recent years. If one draw a pencil-point over the palm or cheek one will notice certain spots of sudden coolness. These are the cold-spots; the heat-spots are less easy to single out. Goldscheider, Blix, and Donaldson have made minute exploration of determinate tracts of skin and found the heat-and cold-spots thick-set and permanently distinct. Between them no temperature-sensation is excited by contact with a pointed cold or hot object. Mechanical and faradic irritation also excites in these points their specific feelings respectively.]

Fig. 26.—The figure marked C P shows the cold-spots, that marked H P the heat-spots, and the middle one the hairs on a certain patch of skin on one of Goldscheider's fingers.

The feeling of temperature is relative to the state of the skin. "In a comfortable room we feel at no part of the body either heat or cold, although different parts of its surface are at different temperatures; the fingers and nose being cooler than the trunk which is covered by clothes, and this, in turn, cooler than the interior of the mouth. The temperature which a given region of the temperature-organ has (as measured by a thermometer) when it feels neither heat nor cold, is its temperature-sensation zero, and is not associated with any one objective temperature; for not only, as we have just seen, does it vary in different parts of the organ, but also on the same part from time to time. Whenever a skin-region has a temperature above its sensation-zero we feel warmth; and vice versa: the sensation is more marked the greater the difference, and the more suddenly it is produced; touching a metallic body, which conducts heat rapidly to or from the skin, causes a more marked hot or cold sensation than touching a worse conductor, as a piece of wood, of the same temperature.

"The change of temperature in the organ may be brought about by changes in the circulatory apparatus (more blood flowing through the skin warms it and less leads to its cooling), or by temperature-changes in gases, liquids, or solids in contact with it. Sometimes we fail to distinguish clearly whether the cause is external or internal; a person coming in from a windy walk often feels a room uncomfortably warm which is not really so; the exercise has accelerated his circulation and tended to warm his skin, but the moving outer air has rapidly conducted off the extra heat; on entering the house the stationary air there does this less quickly, the skin gets hot, and the cause is supposed to be oppressive heat of the room. Hence, frequently, opening windows and sitting in a draught, with its concomitant risks; whereas keeping quiet for five or ten minutes, until the circulation has returned to its normal rate, would attain the same end without danger.

"The acuteness of the temperature-sense is greatest at temperatures within a few degrees of 30° C. (86° F.); at these differences of less than 0.1° C. can be discriminated. As a means of measuring absolute temperatures, however, the skin is very unreliable, on account of the changeability of its sensation-zero. We can localize temperature-sensations much as tactile, but not so accurately."[25]

Muscular Sensation.—The sensation in the muscle itself cannot well be distinguished from that in the tendon or in its insertion. In muscular fatigue the insertions are the places most painfully felt. In muscular rheumatism, however, the whole muscle grows painful; and violent contraction such as that caused by the faradic current, or known as cramp, produces a severe and peculiar pain felt in the whole mass of muscle affected. Sachs also thought that he had demonstrated, both experimentally and anatomically, the existence of special sensory nerve-fibres, distinct from the motor fibres, in the frog's muscle. The latter end in the 'terminal plates,' the former in a network.

Great importance has been attached to the muscular sense as a factor in our perceptions, not only of weight and pressure, but of the space-relations between things generally. Our eyes and our hands, in their explorations of space, move over it and through it. It is usually supposed that without this sense of an intervening motion performed we should not perceive two seen points or two touched points to be separated by an extended interval. I am far from denying the immense participation of experiences of motion in the construction of our space-perceptions. But it is still an open question how our muscles help us in these experiences, whether by their own sensations, or by awakening sensations of motion on our skin, retina, and articular surfaces. The latter seems to me the more probable view, and the reader may be of the same opinion after reading Chapter VI.

Sensibility to Weight.—When we wish to estimate accurately the weight of an object we always, when possible, lift it, and so combine muscular and articular with tactile sensations. By this means we can form much better judgments.

Weber found that whereas ⅓ must be added to a weight resting on the hand for the increase to be felt, the same hand actively 'hefting' the weight could feel an addition of as little as ¹/₁₇. Merkel's recent and very careful experiments, in which the finger pressed down the beam of a balance counterweighted by from 25 to 8020 grams, showed that between 200 and 2000 grams a constant fractional increase of about ¹/₁₃ was felt when there was no movement of the finger, and of about ¹/₁₉ when there was movement. Above and below these limits the discriminative power grew less.

Fig. 27 (after Wundt).

Pain.—The physiology of pain is still an enigma. One might suppose separate afferent fibres with their own end-organs to carry painful impressions to a specific pain-centre. Or one might suppose such a specific centre to be reached by currents of overflow from the other sensory centres when the violence of their inner excitement should have reached a certain pitch. Or again one might suppose a certain extreme degree of inner excitement to produce the feeling of pain in all the centres. It is certain that sensations of every order, which in moderate degrees are rather pleasant than otherwise, become painful when their intensity grows strong. The rate at which the agreeableness and disagreeableness vary with the intensity of a sensation is roughly represented by the dotted curve in Fig. 27. The horizontal line represents the threshold both of sensational and of agreeable sensibility. Below the line is the disagreeble. The continuous curve is that of Weber's law which we learned to know in Fig. 2, p. 18. With the minimal sensation the agreeableness is nil, as the dotted curve shows. It rises at first more slowly than the sensational intensity, then faster; and reaches its maximum before the sensation is near its acme. After its maximum of agreeableness the dotted line rapidly sinks, and soon tumbles below the horizontal into the realm of the disagreeable or painful in which it declines. That all sensations are painful when too strong is a piece of familiar knowledge. Light, sound, odors, the taste of sweet even, cold, heat, and all the skin-sensations, must be moderate to be enjoyed.

The quality of the sensation complicates the question, however, for in some sensations, as bitter, sour, salt, and certain smells, the turning point of the dotted curve must be drawn very near indeed to the beginning of the scale. In the skin the painful quality soon becomes so intense as entirely to overpower the specific quality of the sort of stimulus. Heat, cold, and pressure are indistinguishable when extreme—we only feel the pain. The hypothesis of separate end-organs in the skin receives some corroboration from recent experiments, for both Blix and Goldscheider have found, along with their special heat-and cold spots, also special 'pain-spots' on the skin. Mixed in with these are spots which are quite feelingless. However it may stand with the terminal pain-spots, separate paths of conduction to the brain, for painful and for merely tactile stimulations of the skin, are made probable by certain facts. In the condition termed analgesia, a touch is felt, but the most violent pinch, burn, or electric spark destructive of the tissue will awaken no sensation. This may occur in disease of the cord, by suggestion in hypnotism, or in certain stages of ether and chloroform intoxication. "In rabbits a similar state of things was produced by Schiff, by dividing the gray matter of the cord, leaving the posterior white columns intact. If, on the contrary, the latter were divided and the gray substance left, there was increased sensitiveness to pain, and possibly touch proper was lost. Such experiments make it pretty certain that when afferent impulses reach the spinal cord at any level and there enter its gray matter with the posterior root-fibres, they travel on in different tracts to conscious centres; the tactile ones coming soon out of the gray network and coursing on in a readily conducting white fibre, while the painful ones travel on farther in the gray substance. It is still uncertain if both impulses reach the cord in the same fibres. The gray network conducts nerve-impulses, but not easily; they tend soon to be blocked in it. A feeble (tactile) impulse reaching it by an afferent fibre might only spread a short way and then pass out into a single good conducting fibre in a white column, and proceed to the brain; while a stronger (painful) impulse would radiate farther in the gray matter, and perhaps break out of it by many fibres leading to the brain through the white columns, and so give rise to an incoördinate and ill-localized sensation. That pains are badly localized, and worse the more intense they are, is a well-known fact, which would thus receive an explanation."[26]

Pain also gives rise to ill-coördinated movements of defence. The stronger the pain the more violent the start. Doubtless in low animals pain is almost the only stimulus; and we have preserved the peculiarity in so far that to-day it is the stimulus of our most energetic, though not of our most discriminating, reactions.

Taste, smell, as well as hunger, thirst, nausea, and other so-called 'common' sensations need not be touched on in this book, as almost nothing of psychological interest is known concerning them.