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ORIGIN OF DOMESTIC PLANTS.

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One of the most important questions connected with our subject is: how far we are to regard our cereal grains, our esculent bulbs and roots, and the multiplied tree fruits of our gardens, as artificially modified and improved forms of wild, self-propagating vegetation. The narratives of botanical travellers have often announced the discovery of the original form and habitat of domesticated plants, and scientific journals have described the experiments by which the identity of particular wild and cultivated vegetables has been thought to be established. It is confidently affirmed that maize and the potato—which we must suppose to have been first cultivated at a much later period than the breadstuffs and most other esculent vegetables of Europe and the East—are found wild and self-propagating in Spanish America, though in forms not recognizable by the common observer as identical with the familiar corn and tuber of modern agriculture. It was lately asserted, upon what seemed very strong evidence, that the Aegilops ovata, a plant growing wild in Southern France, had been actually converted into common wheat; but, upon a repetition of the experiments, later observers have declared that the apparent change was only a case of temporary hybridation or fecundation by the pollen of true wheat, and that the grass alleged to be transformed into wheat could not be perpetuated as such from its own seed.

The very great modifications which cultivated plants are constantly undergoing under our eyes, and the numerous varieties and races which spring up among them, certainly countenance the doctrine, that every domesticated vegetable, however dependent upon human care for growth and propagation in its present form, may have been really derived, by a long Succession of changes, from some wild plant not now perhaps much resembling it. [Footnote: What is the possible limit of such changes, we do not know, but they may doubtless be carriad vastly beyond what experience has yet shown to be practicable. Civilized man has experimented little on wild plants, and especially on forest trees. He has indeed improved the fruit, and developed new varieties, of the chestnut, by cultivation, and it is observed that our American forest-tree nuts and berries, such as the butternut and thewild mulberry, become larger and better flavored in a single generation by planting and training. (Bryant, Forest Trees, 1871, pp. 99, 115.) Why should not the industry and ingenuity which have wrought such wonders in our horticulture produce analogous results when applied to the cultivation and amelioration of larger vegetables Might not, for instance, the ivory nut, the fruit of the Phytelephas macrocarpa, possibly be so increased in size as to serve nearly all the purposes of animal ivory now becoming so scarce Might not the various milk-producing trees become, by cultivation, a really important source of nutriment to the inhabitants of warm climates In short, there is room to hope incalculable advantage from the exercise of human skill in the improvement of yet untamed forms of vegetable life.] But it is, in every case, a question of evidence.

The only satisfactory proof that a given wild plant is identical with a given garden or field vegetable, is the test of experiment, the actual growing of the one from the seed of the other, or the conversion of the one into the other by transplantation and change of conditions. [Footnote: The poisonous wild parsnip of New England has been often asserted to be convertible into the common garden parsnip by cultivation, or rather to be the same vegetable growing under different conditions, and it is said to be deprived of its deleterious qualities simply by an increased luxuriance of growth in rich, tilled earth. Wild medicinal plants, so important in the rustic materia medica of New England—such as pennyroyal, for example—are generally much less aromatic and powerful when cultivated in gardens than when self-sown on meagre soils. On the other hand, the cinchona, lately introduced from South America into British India and carefully cultivated there, is found to be richer in quinine than the American tree.]

It is hardly contended that any of the cereals or other plants important as human aliment, or as objects of agricultural industry, exist and propagate themselves uncultivated in the same form and with the same properties as when sown and reared by human art. [Footnote: Some recent observations of Wetzatein are worthy of special notice. "The soil of the Hauran," he remarks, "produces, in its primitive condition, much wild rye, which is not known as a cultivated plant in Syria, and much wild barley and oats. These cereals precisely resemble the corresponding cultivated plants in leaf, ear, size, and height of straw, but their grains are sensibly flatter and poorer in flour."—Reisebericht uber Hauran und die Trachenen, p. 40.

Some of the cereals are, to a certain extent, self-propagating in the soil and climate of California. "VOLUNTEER crops are grown from the seed which falls out in harvesting. Barley has been known to volunteer five crops in succession."—Prayer-Frowd, Six Months in California, p. 189.] In fact, the cases are rare where the identity of a wild with a domesticated plant is considered by the best authorities as conclusively established, and we are warranted in affirming of but few of the latter, as a historically known or experimentally proved fact, that they ever did exist, or could exist, independently of man. [Footnote: This remark is much less applicable to fruit trees than to garden vegetables and the cerealia. The wild orange of Florida, though once considered indigenous, is now generally thought by botanists to be descended from the European orange introduced by the early colonists. On the wild apple trees of Massachusetts see an interesting chapter in Thoreau, Excursions. The fig and the olive are found growing wild in every country where those trees are cultivated The wild fig differs from the domesticated in its habits, its season of fructification, and its insect population, but is, I believe, not specifically distinguishable from the garden fig, though I do not know that it is reclaimable by cultivation. The wild olive, which is so abundant in the Tuscan Maremma, produces good fruit without further care, when thinned out and freed from the shade of other trees, and is particularly suited for grafting. See Salvagnoli, Memorie sulle Maremme, pp. 63–73. The olive is indigenous in Syria and in the Punjaub, and forms vast forests in the Himalayas at from 1,400 to 2,100 feet above the level of the sea.—Cleghorn, Memoir on the Timber procured from the Indus, etc., pp. 8–15. Fraas, Klima und Pfanzenwelt in der Zeit, pp. 35–38, gives, upon the authority of Link and other botanical writers, a lift of the native habitats of most cereals and of many fruits, or at least of localities where those plants are said to be now found wild; but the data do not appear to rest, in general, upon very trustworthy evidence. Theoretically, there can be little doubt that all our cultivated plants are modified forms of spontaneous vegetation, though the connection is not historically shown, nor are we able to say that the originals of some domesticated vegetables may not be now extinct and unrepresented in the existing wild flora. See, on this subject, Humboldt, Ansichten der Natur, i., pp. 208, 209.

The Adams of modern botany and zoology have been put to hard shifts in finding names for the multiplied organisms which the Creator has brought before them, "to see what they would call them;" and naturalists and philosophers have shown much moral courage in setting at naught the law of philology in the coinage of uncouth words to express scientific Ideas. It is much to be wished that some bold neologist would devise English technical equivalents for the German verwildert, run-wild, and veredelt, improved by cultivation.]

Transfer of Vegetable Life.

It belongs to vegetable and animal geography, which are almost sciences of themselves, to point out in detail what man has done to change the distribution of plants and of animated life and to revolutionize the aspect of organic nature; but some of the more important facts bearing on the first branch of this subject may pertinently be introduced here. Most of the cereal grains, the pulse, the edible roots, the tree fruits, and other important forms of esculent vegetation grown in Europe and the United States are believed, and—if the testimony of Pliny and other ancient naturalists is to be depended upon—many of them are historically known, to have originated in the temperate climates of Asia. The agriculture of even so old a country as Egypt has been almost completely revolutionized by the introduction of foreign plants, within the historical period. "With the exception of wheat," says Hehn, "the Nile valley now yields only new products, cotton, rice, sugar, indigo, sorghum, dates," being all unknown to its most ancient rural husbandry. [Footnote: On these points see the learned work of Hehn, Kultur. Pflanzen und Thiere in ihrem Uebergang aus Asien, 1870. On the migration of plants generally, see Lyell, Principles of Geology, 10th ed., vol. ii., c.] The wine grape has been thought to be truly indigenous only in the regions bordering on the eastern end of the Black Sea, where it now, particularly on the banks of the Rion, the ancient Phasis, propagates itself spontaneously, and grows with unexampled luxuriance. [Footnote: The vine-wood planks of the ancient great door of the cathedral at Ravenna, which measured thirteen feet in length by a foot and a quarter in width, are traditionally said to have boon brought from the Black Sea, by way of Constantinople, about the eleventh or twelfth century. Vines of such dimension are now very rarely found in any other part of the East, and, though I have taken some pains on the subject, I never found in Syria or in Turkey a vine stock exceeding six inches in diameter, bark excluded. Schulz, however, saw at Beitschin, near Ptolemais, a vine measuring eighteen inches in diameter. Strabo speaks of vine-stocks in Margiana (Khorasan) of such dimension that two men, with outstretched arms, could scarcely embrace them. See Strabo, ed. Casaubon, pp. 78, 516, 826. Statues of vine wood are mentioned by ancient writers. Very large vine-stems are not common in Italy, but the vine-wood panels of the door of the chapter-hall of the church of St. John at Saluzzo are not less than ten inches in width, and I observed not long since, in a garden at Pie di Mulera, a vine stock with a circumference of thirty inches.] But some species of the vine seem native to Europe, and many varieties of grape have been too long known as common to every part of the United States to admit of the supposition that they were introduced by European colonists. [Footnote: The Northmen who—as I think it has been indisputably established by Professor Rafn of Copenhagen—visited the coast of Massachusetts about theyear 1000, found grapes growing there in profusion, and the wild vine still flourishes in great variety and abundance in the southeastern counties of that State. The townships in the vicinity of the Dighton rock, supposed by many—with whom, however, I am sorry I cannot agree—to bear a Scandinavian inscription, abound in wild vines. According to Laudonniere, Histoire Notable de la Florida, reprint, Paris, 1853, p 5, the French navigators in 1562 found in that peninsula "wild vines which climb the trees and produce good grapes."]

The Earth as Modified by Human Action

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