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In most mammals, the cauda epididymis is cooler than the testes [34], facilitating its sperm storage function. Increasing cauda temperature disrupts absorptive and secretory functions, changes the composition (ions and proteins) of the cauda fluid, and increases (approximately threefold) the rate of sperm passage through the cauda [34]. Consequently, the number of sperm in the first ejaculate declines, with an even more dramatic decline in sperm number in successive ejaculates. In addition, increased temperature seems to hasten sperm maturation [34].