Читать книгу Creative Evolution - Henri Bergson - Страница 7

To sum up, if the accidental variations that bring about evolution are insensible variations, some good genius must be appealed to—the genius of the future species—in order to preserve and accumulate these variations, for selection will not look after this. If, on the other hand, the accidental variations are sudden, then, for the previous function to go on or for a new function to take its place, all the changes that have happened together must be complementary. So we have to fall back on the good genius again, this time to obtain the convergence of simultaneous changes, as before to be assured of the continuity of direction of successive variations. But in neither case can parallel development of the same complex structures on independent lines of evolution be due to a mere accumulation of accidental variations. So we come to the second of the two great hypotheses we have to examine. Suppose the variations are due, not to accidental and inner causes, but to the direct influence of outer circumstances. Let us see what line we should have to take, on this hypothesis, to account for the resemblance of eye-structure in two series that are independent of each other from the phylogenetic point of view.

Though molluscs and vertebrates have evolved separately, both have remained exposed to the influence of light. And light is a physical cause bringing forth certain definite effects. Acting in a continuous way, it has been able to produce a continuous variation in a constant direction. Of course it is unlikely that the eye of the vertebrate and that of the mollusc have been built up by a series of variations due to simple chance. Admitting even that light enters into the case as an instrument of selection, in order to allow only useful variations to persist, there is no possibility that the play of chance, even thus supervised from without, should bring about in both cases the same juxtaposition of elements coördinated in the same way. But it would be different supposing that light acted directly on the organized matter so as to change its structure and somehow adapt this structure to its own form. The resemblance of the two effects would then be explained by the identity of the cause. The more and more complex eye would be something like the deeper and deeper imprint of light on a matter which, being organized, possesses a special aptitude for receiving it.

But can an organic structure be likened to an imprint? We have already called attention to the ambiguity of the term "adaptation." The gradual complication of a form which is being better and better adapted to the mold of outward circumstances is one thing, the increasingly complex structure of an instrument which derives more and more advantage from these circumstances is another. In the former case, the matter merely receives an imprint; in the second, it reacts positively, it solves a problem. Obviously it is this second sense of the word "adapt" that is used when one says that the eye has become better and better adapted to the influence of light. But one passes more or less unconsciously from this sense to the other, and a purely mechanistic biology will strive to make the passive adaptation of an inert matter, which submits to the influence of its environment, mean the same as the active adaptation of an organism which derives from this influence an advantage it can appropriate. It must be owned, indeed, that Nature herself appears to invite our mind to confuse these two kinds of adaptation, for she usually begins by a passive adaptation where, later on, she will build up a mechanism for active response. Thus, in the case before us, it is unquestionable that the first rudiment of the eye is found in the pigment-spot of the lower organisms; this spot may indeed have been produced physically, by the mere action of light, and there are a great number of intermediaries between the simple spot of pigment and a complicated eye like that of the vertebrates.—But, from the fact that we pass from one thing to another by degrees, it does not follow that the two things are of the same nature. From the fact that an orator falls in, at first, with the passions of his audience in order to make himself master of them, it will not be concluded that to follow is the same as to lead. Now, living matter seems to have no other means of turning circumstances to good account than by adapting itself to them passively at the outset. Where it has to direct a movement, it begins by adopting it. Life proceeds by insinuation. The intermediate degrees between a pigment-spot and an eye are nothing to the point: however numerous the degrees, there will still be the same interval between the pigment-spot and the eye as between a photograph and a photographic apparatus. Certainly the photograph has been gradually turned into a photographic apparatus; but could light alone, a physical force, ever have provoked this change, and converted an impression left by it into a machine capable of using it?

It may be claimed that considerations of utility are out of place here; that the eye is not made to see, but that we see because we have eyes; that the organ is what it is, and "utility" is a word by which we designate the functional effects of the structure. But when I say that the eye "makes use of" light, I do not merely mean that the eye is capable of seeing; I allude to the very precise relations that exist between this organ and the apparatus of locomotion. The retina of vertebrates is prolonged in an optic nerve, which, again, is continued by cerebral centres connected with motor mechanisms. Our eye makes use of light in that it enables us to utilize, by movements of reaction, the objects that we see to be advantageous, and to avoid those which we see to be injurious. Now, of course, as light may have produced a pigment-spot by physical means, so it can physically determine the movements of certain organisms; ciliated Infusoria, for instance, react to light. But no one would hold that the influence of light has physically caused the formation of a nervous system, of a muscular system, of an osseous system, all things which are continuous with the apparatus of vision in vertebrate animals. The truth is, when one speaks of the gradual formation of the eye, and, still more, when one takes into account all that is inseparably connected with it, one brings in something entirely different from the direct action of light. One implicitly attributes to organized matter a certain capacity sui generis, the mysterious power of building up very complicated machines to utilize the simple excitation that it undergoes.

But this is just what is claimed to be unnecessary. Physics and chemistry are said to give us the key to everything. Eimer's great work is instructive in this respect. It is well known what persevering effort this biologist has devoted to demonstrating that transformation is brought about by the influence of the external on the internal, continuously exerted in the same direction, and not, as Darwin held, by accidental variations. His theory rests on observations of the highest interest, of which the starting-point was the study of the course followed by the color variation of the skin in certain lizards. Before this, the already old experiments of Dorfmeister had shown that the same chrysalis, according as it was submitted to cold or heat, gave rise to very different butterflies, which had long been regarded as independent species, Vanessa levana and Vanessa prorsa: an intermediate temperature produces an intermediate form. We might class with these facts the important transformations observed in a little crustacean, Artemia salina, when the salt of the water it lives in is increased or diminished.[32] In these various experiments the external agent seems to act as a cause of transformation. But what does the word "cause" mean here? Without undertaking an exhaustive analysis of the idea of causality, we will merely remark that three very different meanings of this term are commonly confused. A cause may act by impelling, releasing, or unwinding. The billiard-ball, that strikes another, determines its movement by impelling. The spark that explodes the powder acts by releasing. The gradual relaxing of the spring, that makes the phonograph turn, unwinds the melody inscribed on the cylinder: if the melody which is played be the effect, and the relaxing of the spring the cause, we must say that the cause acts by unwinding. What distinguishes these three cases from each other is the greater or less solidarity between the cause and the effect. In the first, the quantity and quality of the effect vary with the quantity and quality of the cause. In the second, neither quality nor quantity of the effect varies with quality and quantity of the cause: the effect is invariable. In the third, the quantity of the effect depends on the quantity of the cause, but the cause does not influence the quality of the effect: the longer the cylinder turns by the action of the spring, the more of the melody I shall hear, but the nature of the melody, or of the part heard, does not depend on the action of the spring. Only in the first case, really, does cause explain effect; in the others the effect is more or less given in advance, and the antecedent invoked is—in different degrees, of course—its occasion rather than its cause. Now, in saying that the saltness of the water is the cause of the transformations of Artemia, or that the degree of temperature determines the color and marks of the wings which a certain chrysalis will assume on becoming a butterfly, is the word "cause" used in the first sense? Obviously not: causality has here an intermediary sense between those of unwinding and releasing. Such, indeed, seems to be Eimer's own meaning when he speaks of the "kaleidoscopic" character of the variation,[33] or when he says that the variation of organized matter works in a definite way, just as inorganic matter crystallizes in definite directions.[34] And it may be granted, perhaps, that the process is a merely physical and chemical one in the case of the color-changes of the skin. But if this sort of explanation is extended to the case of the gradual formation of the eye of the vertebrate, for instance, it must be supposed that the physico-chemistry of living bodies is such that the influence of light has caused the organism to construct a progressive series of visual apparatus, all extremely complex, yet all capable of seeing, and of seeing better and better.[35] What more could the most confirmed finalist say, in order to mark out so exceptional a physico-chemistry? And will not the position of a mechanistic philosophy become still more difficult, when it is pointed out to it that the egg of a mollusc cannot have the same chemical composition as that of a vertebrate, that the organic substance which evolved toward the first of these two forms could not have been chemically identical with that of the substance which went in the other direction, and that, nevertheless, under the influence of light, the same organ has been constructed in the one case as in the other?

The more we reflect upon it, the more we shall see that this production of the same effect by two different accumulations of an enormous number of small causes is contrary to the principles of mechanistic philosophy. We have concentrated the full force of our discussion upon an example drawn from phylogenesis. But ontogenesis would have furnished us with facts no less cogent. Every moment, right before our eyes, nature arrives at identical results, in sometimes neighboring species, by entirely different embryogenic processes. Observations of "heteroblastia" have multiplied in late years,[36] and it has been necessary to reject the almost classical theory of the specificity of embryonic gills. Still keeping to our comparison between the eye of vertebrates and that of molluscs, we may point out that the retina of the vertebrate is produced by an expansion in the rudimentary brain of the young embryo. It is a regular nervous centre which has moved toward the periphery. In the mollusc, on the contrary, the retina is derived from the ectoderm directly, and not indirectly by means of the embryonic encephalon. Quite different, therefore, are the evolutionary processes which lead, in man and in the Pecten, to the development of a like retina. But, without going so far as to compare two organisms so distant from each other, we might reach the same conclusion simply by looking at certain very curious facts of regeneration in one and the same organism. If the crystalline lens of a Triton be removed, it is regenerated by the iris.[37] Now, the original lens was built out of the ectoderm, while the iris is of mesodermic origin. What is more, in the Salamandra maculata, if the lens be removed and the iris left, the regeneration of the lens takes place at the upper part of the iris; but if this upper part of the iris itself be taken away, the regeneration takes place in the inner or retinal layer of the remaining region.[38] Thus, parts differently situated, differently constituted, meant normally for different functions, are capable of performing the same duties and even of manufacturing, when necessary, the same pieces of the machine. Here we have, indeed, the same effect obtained by different combinations of causes.

Whether we will or no, we must appeal to some inner directing principle in order to account for this convergence of effects. Such convergence does not appear possible in the Darwinian, and especially the neo-Darwinian, theory of insensible accidental variations, nor in the hypothesis of sudden accidental variations, nor even in the theory that assigns definite directions to the evolution of the various organs by a kind of mechanical composition of the external with the internal forces. So we come to the only one of the present forms of evolution which remains for us to mention, viz., neo-Lamarckism.

It is well known that Lamarck attributed to the living being the power of varying by use or disuse of its organs, and also of passing on the variation so acquired to its descendants. A certain number of biologists hold a doctrine of this kind to-day. The variation that results in a new species is not, they believe, merely an accidental variation inherent in the germ itself, nor is it governed by a determinism sui generis which develops definite characters in a definite direction, apart from every consideration of utility. It springs from the very effort of the living being to adapt itself to the circumstances of its existence. The effort may indeed be only the mechanical exercise of certain organs, mechanically elicited by the pressure of external circumstances. But it may also imply consciousness and will, and it is in this sense that it appears to be understood by one of the most eminent representatives of the doctrine, the American naturalist Cope.[39] Neo-Lamarckism is therefore, of all the later forms of evolutionism, the only one capable of admitting an internal and psychological principle of development, although it is not bound to do so. And it is also the only evolutionism that seems to us to account for the building up of identical complex organs on independent lines of development. For it is quite conceivable that the same effort to turn the same circumstances to good account might have the same result, especially if the problem put by the circumstances is such as to admit of only one solution. But the question remains, whether the term "effort" must not then be taken in a deeper sense, a sense even more psychological than any neo-Lamarckian supposes.

For a mere variation of size is one thing, and a change of form is another. That an organ can be strengthened and grow by exercise, nobody will deny. But it is a long way from that to the progressive development of an eye like that of the molluscs and of the vertebrates. If this development be ascribed to the influence of light, long continued but passively received, we fall back on the theory we have just criticized. If, on the other hand, an internal activity is appealed to, then it must be something quite different from what we usually call an effort, for never has an effort been known to produce the slightest complication of an organ, and yet an enormous number of complications, all admirably coördinated, have been necessary to pass from the pigment-spot of the Infusorian to the eye of the vertebrate. But, even if we accept this notion of the evolutionary process in the case of animals, how can we apply it to plants? Here, variations of form do not seem to imply, nor always to lead to, functional changes; and even if the cause of the variation is of a psychological nature, we can hardly call it an effort, unless we give a very unusual extension to the meaning of the word. The truth is, it is necessary to dig beneath the effort itself and look for a deeper cause.

This is especially necessary, we believe, if we wish to get at a cause of regular hereditary variations. We are not going to enter here into the controversies over the transmissibility of acquired characters; still less do we wish to take too definite a side on this question, which is not within our province. But we cannot remain completely indifferent to it. Nowhere is it clearer that philosophers can not to-day content themselves with vague generalities, but must follow the scientists in experimental detail and discuss the results with them. If Spencer had begun by putting to himself the question of the hereditability of acquired characters, his evolutionism would no doubt have taken an altogether different form. If (as seems probable to us) a habit contracted by the individual were transmitted to its descendants only in very exceptional cases, all the Spencerian psychology would need remaking, and a large part of Spencer's philosophy would fall to pieces. Let us say, then, how the problem seems to us to present itself, and in what direction an attempt might be made to solve it.

After having been affirmed as a dogma, the transmissibility of acquired characters has been no less dogmatically denied, for reasons drawn a priori from the supposed nature of germinal cells. It is well known how Weismann was led, by his hypothesis of the continuity of the germ-plasm, to regard the germinal cells—ova and spermatozoa—as almost independent of the somatic cells. Starting from this, it has been claimed, and is still claimed by many, that the hereditary transmission of an acquired character is inconceivable. But if, perchance, experiment should show that acquired characters are transmissible, it would prove thereby that the germ-plasm is not so independent of the somatic envelope as has been contended, and the transmissibility of acquired characters would become ipso facto conceivable; which amounts to saying that conceivability and inconceivability have nothing to do with the case, and that experience alone must settle the matter. But it is just here that the difficulty begins. The acquired characters we are speaking of are generally habits or the effects of habit, and at the root of most habits there is a natural disposition. So that one can always ask whether it is really the habit acquired by the soma of the individual that is transmitted, or whether it is not rather a natural aptitude, which existed prior to the habit. This aptitude would have remained inherent in the germ-plasm which the individual bears within him, as it was in the individual himself and consequently in the germ whence he sprang. Thus, for instance, there is no proof that the mole has become blind because it has formed the habit of living underground; it is perhaps because its eyes were becoming atrophied that it condemned itself to a life underground.[40] If this is the case, the tendency to lose the power of vision has been transmitted from germ to germ without anything being acquired or lost by the soma of the mole itself. From the fact that the son of a fencing-master has become a good fencer much more quickly than his father, we cannot infer that the habit of the parent has been transmitted to the child; for certain natural dispositions in course of growth may have passed from the plasma engendering the father to the plasma engendering the son, may have grown on the way by the effect of the primitive impetus, and thus assured to the son a greater suppleness than the father had, without troubling, so to speak, about what the father did. So of many examples drawn from the progressive domestication of animals: it is hard to say whether it is the acquired habit that is transmitted or only a certain natural tendency—that, indeed, which has caused such and such a particular species or certain of its representatives to be specially chosen for domestication. The truth is, when every doubtful case, every fact open to more than one interpretation, has been eliminated, there remains hardly a single unquestionable example of acquired and transmitted peculiarities, beyond the famous experiments of Brown-Séquard, repeated and confirmed by other physiologists.[41] By cutting the spinal cord or the sciatic nerve of guinea-pigs, Brown-Séquard brought about an epileptic state which was transmitted to the descendants. Lesions of the same sciatic nerve, of the restiform body, etc., provoked various troubles in the guinea-pig which its progeny inherited sometimes in a quite different form: exophthalmia, loss of toes, etc. But it is not demonstrated that in these different cases of hereditary transmission there had been a real influence of the soma of the animal on its germ-plasm. Weismann at once objected that the operations of Brown-Séquard might have introduced certain special microbes into the body of the guinea-pig, which had found their means of nutrition in the nervous tissues and transmitted the malady by penetrating into the sexual elements.[42] This objection has been answered by Brown-Séquard himself;[43] but a more plausible one might be raised. Some experiments of Voisin and Peron have shown that fits of epilepsy are followed by the elimination of a toxic body which, when injected into animals,[44] is capable of producing convulsive symptoms. Perhaps the trophic disorders following the nerve lesions made by Brown-Séquard correspond to the formation of precisely this convulsion-causing poison. If so, the toxin passed from the guinea-pig to its spermatozoon or ovum, and caused in the development of the embryo a general disturbance, which, however, had no visible effects except at one point or another of the organism when developed. In that case, what occurred would have been somewhat the same as in the experiments of Charrin, Delamare, and Moussu, where guinea-pigs in gestation, whose liver or kidney was injured, transmitted the lesion to their progeny, simply because the injury to the mother's organ had given rise to specific "cytotoxins" which acted on the corresponding organ of the foetus.[45] It is true that, in these experiments, as in a former observation of the same physiologists,[46] it was the already formed foetus that was influenced by the toxins. But other researches of Charrin have resulted in showing that the same effect may be produced, by an analogous process, on the spermatozoa and the ova.[47] To conclude, then: the inheritance of an acquired peculiarity in the experiments of Brown-Séquard can be explained by the effect of a toxin on the germ. The lesion, however well localized it seems, is transmitted by the same process as, for instance, the taint of alcoholism. But may it not be the same in the case of every acquired peculiarity that has become hereditary?

There is, indeed, one point on which both those who affirm and those who deny the transmissibility of acquired characters are agreed, namely, that certain influences, such as that of alcohol, can affect at the same time both the living being and the germ-plasm it contains. In such case, there is inheritance of a defect, and the result is as if the soma of the parent had acted on the germ-plasm, although in reality soma and plasma have simply both suffered the action of the same cause. Now, suppose that the soma can influence the germ-plasm, as those believe who hold that acquired characters are transmissible. Is not the most natural hypothesis to suppose that things happen in this second case as in the first, and that the direct effect of the influence of the soma is a general alteration of the germ-plasm? If this is the case, it is by exception, and in some sort by accident, that the modification of the descendant is the same as that of the parent. It is like the hereditability of the alcoholic taint: it passes from father to children, but it may take a different form in each child, and in none of them be like what it was in the father. Let the letter C represent the change in the plasm, C being either positive or negative, that is to say, showing either the gain or loss of certain substances. The effect will not be an exact reproduction of the cause, nor will the change in the germ-plasm, provoked by a certain modification of a certain part of the soma, determine a similar modification of the corresponding part of the new organism in process of formation, unless all the other nascent parts of this organism enjoy a kind of immunity as regards C: the same part will then undergo alteration in the new organism, because it happens that the development of this part is alone subject to the new influence. And, even then, the part might be altered in an entirely different way from that in which the corresponding part was altered in the generating organism.

We should propose, then, to introduce a distinction between the hereditability of deviation and that of character. An individual which acquires a new character thereby deviates from the form it previously had, which form the germs, or oftener the half-germs, it contains would have reproduced in their development. If this modification does not involve the production of substances capable of changing the germ-plasm, or does not so affect nutrition as to deprive the germ-plasm of certain of its elements, it will have no effect on the offspring of the individual. This is probably the case as a rule. If, on the contrary, it has some effect, this is likely to be due to a chemical change which it has induced in the germ-plasm. This chemical change might, by exception, bring about the original modification again in the organism which the germ is about to develop, but there are as many and more chances that it will do something else. In this latter case, the generated organism will perhaps deviate from the normal type as much as the generating organism, but it will do so differently. It will have inherited deviation and not character. In general, therefore, the habits formed by an individual have probably no echo in its offspring; and when they have, the modification in the descendants may have no visible likeness to the original one. Such, at least, is the hypothesis which seems to us most likely. In any case, in default of proof to the contrary, and so long as the decisive experiments called for by an eminent biologist[48] have not been made, we must keep to the actual results of observation. Now, even if we take the most favorable view of the theory of the transmissibility of acquired characters, and assume that the ostensible acquired character is not, in most cases, the more or less tardy development of an innate character, facts show us that hereditary transmission is the exception and not the rule. How, then, shall we expect it to develop an organ such as the eye? When we think of the enormous number of variations, all in the same direction, that we must suppose to be accumulated before the passage from the pigment-spot of the Infusorian to the eye of the mollusc and of the vertebrate is possible, we do not see how heredity, as we observe it, could ever have determined this piling-up of differences, even supposing that individual efforts could have produced each of them singly. That is to say that neo-Lamarckism is no more able than any other form of evolutionism to solve the problem.

In thus submitting the various present forms of evolutionism to a common test, in showing that they all strike against the same insurmountable difficulty, we have in no wise the intention of rejecting them altogether. On the contrary, each of them, being supported by a considerable number of facts, must be true in its way. Each of them must correspond to a certain aspect of the process of evolution. Perhaps even it is necessary that a theory should restrict itself exclusively to a particular point of view, in order to remain scientific, i.e. to give a precise direction to researches into detail. But the reality of which each of these theories takes a partial view must transcend them all. And this reality is the special object of philosophy, which is not constrained to scientific precision because it contemplates no practical application. Let us therefore indicate in a word or two the positive contribution that each of the three present forms of evolutionism seems to us to make toward the solution of the problem, what each of them leaves out, and on what point this threefold effort should, in our opinion, converge in order to obtain a more comprehensive, although thereby of necessity a less definite, idea of the evolutionary process.

The neo-Darwinians are probably right, we believe, when they teach that the essential causes of variation are the differences inherent in the germ borne by the individual, and not the experiences or behavior of the individual in the course of his career. Where we fail to follow these biologists, is in regarding the differences inherent in the germ as purely accidental and individual. We cannot help believing that these differences are the development of an impulsion which passes from germ to germ across the individuals, that they are therefore not pure accidents, and that they might well appear at the same time, in the same form, in all the representatives of the same species, or at least in a certain number of them. Already, in fact, the theory of mutations is modifying Darwinism profoundly on this point. It asserts that at a given moment, after a long period, the entire species is beset with a tendency to change. The tendency to change, therefore, is not accidental. True, the change itself would be accidental, since the mutation works, according to De Vries, in different directions in the different representatives of the species. But, first we must see if the theory is confirmed by many other vegetable species (De Vries has verified it only by the [OE]nothera Lamarckiana),[49] and then there is the possibility, as we shall explain further on, that the part played by chance is much greater in the variation of plants than in that of animals, because, in the vegetable world, function does not depend so strictly on form. Be that as it may, the neo-Darwinians are inclined to admit that the periods of mutation are determinate. The direction of the mutation may therefore be so as well, at least in animals, and to the extent we shall have to indicate.

We thus arrive at a hypothesis like Eimer's, according to which the variations of different characters continue from generation to generation in definite directions. This hypothesis seems plausible to us, within the limits in which Eimer himself retains it. Of course, the evolution of the organic world cannot be predetermined as a whole. We claim, on the contrary, that the spontaneity of life is manifested by a continual creation of new forms succeeding others. But this indetermination cannot be complete; it must leave a certain part to determination. An organ like the eye, for example, must have been formed by just a continual changing in a definite direction. Indeed, we do not see how otherwise to explain the likeness of structure of the eye in species that have not the same history. Where we differ from Eimer is in his claim that combinations of physical and chemical causes are enough to secure the result. We have tried to prove, on the contrary, by the example of the eye, that if there is "orthogenesis" here, a psychological cause intervenes.

Certain neo-Lamarckians do indeed resort to a cause of a psychological nature. There, to our thinking, is one of the most solid positions of neo-Lamarckism. But if this cause is nothing but the conscious effort of the individual, it cannot operate in more than a restricted number of cases—at most in the animal world, and not at all in the vegetable kingdom. Even in animals, it will act only on points which are under the direct or indirect control of the will. And even where it does act, it is not clear how it could compass a change so profound as an increase of complexity: at most this would be conceivable if the acquired characters were regularly transmitted so as to be added together; but this transmission seems to be the exception rather than the rule. A hereditary change in a definite direction, which continues to accumulate and add to itself so as to build up a more and more complex machine, must certainly be related to some sort of effort, but to an effort of far greater depth than the individual effort, far more independent of circumstances, an effort common to most representatives of the same species, inherent in the germs they bear rather than in their substance alone, an effort thereby assured of being passed on to their descendants.

So we come back, by a somewhat roundabout way, to the idea we started from, that of an original impetus of life, passing from one generation of germs to the following generation of germs through the developed organisms which bridge the interval between the generations. This impetus, sustained right along the lines of evolution among which it gets divided, is the fundamental cause of variations, at least of those that are regularly passed on, that accumulate and create new species. In general, when species have begun to diverge from a common stock, they accentuate their divergence as they progress in their evolution. Yet, in certain definite points, they may evolve identically; in fact, they must do so if the hypothesis of a common impetus be accepted. This is just what we shall have to show now in a more precise way, by the same example we have chosen, the formation of the eye in molluscs and vertebrates. The idea of an "original impetus," moreover, will thus be made clearer.

Two points are equally striking in an organ like the eye: the complexity of its structure and the simplicity of its function. The eye is composed of distinct parts, such as the sclerotic, the cornea, the retina, the crystalline lens, etc. In each of these parts the detail is infinite. The retina alone comprises three layers of nervous elements—multipolar cells, bipolar cells, visual cells—each of which has its individuality and is undoubtedly a very complicated organism: so complicated, indeed, is the retinal membrane in its intimate structure, that no simple description can give an adequate idea of it. The mechanism of the eye is, in short, composed of an infinity of mechanisms, all of extreme complexity. Yet vision is one simple fact. As soon as the eye opens, the visual act is effected. Just because the act is simple, the slightest negligence on the part of nature in the building of the infinitely complex machine would have made vision impossible. This contrast between the complexity of the organ and the unity of the function is what gives us pause.

A mechanistic theory is one which means to show us the gradual building-up of the machine under the influence of external circumstances intervening either directly by action on the tissues or indirectly by the selection of better-adapted ones. But, whatever form this theory may take, supposing it avails at all to explain the detail of the parts, it throws no light on their correlation.

Then comes the doctrine of finality, which says that the parts have been brought together on a preconceived plan with a view to a certain end. In this it likens the labor of nature to that of the workman, who also proceeds by the assemblage of parts with a view to the realization of an idea or the imitation of a model. Mechanism, here, reproaches finalism with its anthropomorphic character, and rightly. But it fails to see that itself proceeds according to this method—somewhat mutilated! True, it has got rid of the end pursued or the ideal model. But it also holds that nature has worked like a human being by bringing parts together, while a mere glance at the development of an embryo shows that life goes to work in a very different way. Life does not proceed by the association and addition of elements, but by dissociation and division.

We must get beyond both points of view, both mechanism and finalism being, at bottom, only standpoints to which the human mind has been led by considering the work of man. But in what direction can we go beyond them? We have said that in analyzing the structure of an organ, we can go on decomposing for ever, although the function of the whole is a simple thing. This contrast between the infinite complexity of the organ and the extreme simplicity of the function is what should open our eyes.

In general, when the same object appears in one aspect and in another as infinitely complex, the two aspects have by no means the same importance, or rather the same degree of reality. In such cases, the simplicity belongs to the object itself, and the infinite complexity to the views we take in turning around it, to the symbols by which our senses or intellect represent it to us, or, more generally, to elements of a different order, with which we try to imitate it artificially, but with which it remains incommensurable, being of a different nature. An artist of genius has painted a figure on his canvas. We can imitate his picture with many-colored squares of mosaic. And we shall reproduce the curves and shades of the model so much the better as our squares are smaller, more numerous and more varied in tone. But an infinity of elements infinitely small, presenting an infinity of shades, would be necessary to obtain the exact equivalent of the figure that the artist has conceived as a simple thing, which he has wished to transport as a whole to the canvas, and which is the more complete the more it strikes us as the projection of an indivisible intuition. Now, suppose our eyes so made that they cannot help seeing in the work of the master a mosaic effect. Or suppose our intellect so made that it cannot explain the appearance of the figure on the canvas except as a work of mosaic. We should then be able to speak simply of a collection of little squares, and we should be under the mechanistic hypothesis. We might add that, beside the materiality of the collection, there must be a plan on which the artist worked; and then we should be expressing ourselves as finalists. But in neither case should we have got at the real process, for there are no squares brought together. It is the picture, i.e. the simple act, projected on the canvas, which, by the mere fact of entering into our perception, is decomposed before our eyes into thousands and thousands of little squares which present, as recomposed, a wonderful arrangement. So the eye, with its marvelous complexity of structure, may be only the simple act of vision, divided for us into a mosaic of cells, whose order seems marvelous to us because we have conceived the whole as an assemblage.

If I raise my hand from A to B, this movement appears to me under two aspects at once. Felt from within, it is a simple, indivisible act. Perceived from without, it is the course of a certain curve, AB. In this curve I can distinguish as many positions as I please, and the line itself might be defined as a certain mutual coördination of these positions. But the positions, infinite in number, and the order in which they are connected, have sprung automatically from the indivisible act by which my hand has gone from A to B. Mechanism, here, would consist in seeing only the positions. Finalism would take their order into account. But both mechanism and finalism would leave on one side the movement, which is reality itself. In one sense, the movement is more than the positions and than their order; for it is sufficient to make it in its indivisible simplicity to secure that the infinity of the successive positions as also their order be given at once—with something else which is neither order nor position but which is essential, the mobility. But, in another sense, the movement is less than the series of positions and their connecting order; for, to arrange points in a certain order, it is necessary first to conceive the order and then to realize it with points, there must be the work of assemblage and there must be intelligence, whereas the simple movement of the hand contains nothing of either. It is not intelligent, in the human sense of the word, and it is not an assemblage, for it is not made up of elements. Just so with the relation of the eye to vision. There is in vision more than the component cells of the eye and their mutual coördination: in this sense, neither mechanism nor finalism go far enough. But, in another sense, mechanism and finalism both go too far, for they attribute to Nature the most formidable of the labors of Hercules in holding that she has exalted to the simple act of vision an infinity of infinitely complex elements, whereas Nature has had no more trouble in making an eye than I have in lifting my hand. Nature's simple act has divided itself automatically into an infinity of elements which are then found to be coördinated to one idea, just as the movement of my hand has dropped an infinity of points which are then found to satisfy one equation.

We find it very hard to see things in that light, because we cannot help conceiving organization as manufacturing. But it is one thing to manufacture, and quite another to organize. Manufacturing is peculiar to man. It consists in assembling parts of matter which we have cut out in such manner that we can fit them together and obtain from them a common action. The parts are arranged, so to speak, around the action as an ideal centre. To manufacture, therefore, is to work from the periphery to the centre, or, as the philosophers say, from the many to the one. Organization, on the contrary, works from the centre to the periphery. It begins in a point that is almost a mathematical point, and spreads around this point by concentric waves which go on enlarging. The work of manufacturing is the more effective, the greater the quantity of matter dealt with. It proceeds by concentration and compression. The organizing act, on the contrary, has something explosive about it: it needs at the beginning the smallest possible place, a minimum of matter, as if the organizing forces only entered space reluctantly. The spermatozoon, which sets in motion the evolutionary process of the embryonic life, is one of the smallest cells of the organism; and it is only a small part of the spermatozoon which really takes part in the operation.

But these are only superficial differences. Digging beneath them, we think, a deeper difference would be found.

A manufactured thing delineates exactly the form of the work of manufacturing it. I mean that the manufacturer finds in his product exactly what he has put into it. If he is going to make a machine, he cuts out its pieces one by one and then puts them together: the machine, when made, will show both the pieces and their assemblage. The whole of the result represents the whole of the work; and to each part of the work corresponds a part of the result.

Now I recognize that positive science can and should proceed as if organization was like making a machine. Only so will it have any hold on organized bodies. For its object is not to show us the essence of things, but to furnish us with the best means of acting on them. Physics and chemistry are well advanced sciences, and living matter lends itself to our action only so far as we can treat it by the processes of our physics and chemistry. Organization can therefore only be studied scientifically if the organized body has first been likened to a machine. The cells will be the pieces of the machine, the organism their assemblage, and the elementary labors which have organized the parts will be regarded as the real elements of the labor which has organized the whole. This is the standpoint of science. Quite different, in our opinion, is that of philosophy.

For us, the whole of an organized machine may, strictly speaking, represent the whole of the organizing work (this is, however, only approximately true), yet the parts of the machine do not correspond to parts of the work, because the materiality of this machine does not represent a sum of means employed, but a sum of obstacles avoided: it is a negation rather than a positive reality. So, as we have shown in a former study, vision is a power which should attain by right an infinity of things inaccessible to our eyes. But such a vision would not be continued into action; it might suit a phantom, but not a living being. The vision of a living being is an effective vision, limited to objects on which the being can act: it is a vision that is canalized, and the visual apparatus simply symbolizes the work of canalizing. Therefore the creation of the visual apparatus is no more explained by the assembling of its anatomic elements than the digging of a canal could be explained by the heaping up of the earth which might have formed its banks. A mechanistic theory would maintain that the earth had been brought cart-load by cart-load; finalism would add that it had not been dumped down at random, that the carters had followed a plan. But both theories would be mistaken, for the canal has been made in another way.

With greater precision, we may compare the process by which nature constructs an eye to the simple act by which we raise the hand. But we supposed at first that the hand met with no resistance. Let us now imagine that, instead of moving in air, the hand has to pass through iron filings which are compressed and offer resistance to it in proportion as it goes forward. At a certain moment the hand will have exhausted its effort, and, at this very moment, the filings will be massed and coördinated in a certain definite form, to wit, that of the hand that is stopped and of a part of the arm. Now, suppose that the hand and arm are invisible. Lookers-on will seek the reason of the arrangement in the filings themselves and in forces within the mass. Some will account for the position of each filing by the action exerted upon it by the neighboring filings: these are the mechanists. Others will prefer to think that a plan of the whole has presided over the detail of these elementary actions: they are the finalists. But the truth is that there has been merely one indivisible act, that of the hand passing through the filings: the inexhaustible detail of the movement of the grains, as well as the order of their final arrangement, expresses negatively, in a way, this undivided movement, being the unitary form of a resistance, and not a synthesis of positive elementary actions. For this reason, if the arrangement of the grains is termed an "effect" and the movement of the hand a "cause," it may indeed be said that the whole of the effect is explained by the whole of the cause, but to parts of the cause parts of the effect will in no wise correspond. In other words, neither mechanism nor finalism will here be in place, and we must resort to an explanation of a different kind. Now, in the hypothesis we propose, the relation of vision to the visual apparatus would be very nearly that of the hand to the iron filings that follow, canalize and limit its motion.

The greater the effort of the hand, the farther it will go into the filings. But at whatever point it stops, instantaneously and automatically the filings coördinate and find their equilibrium. So with vision and its organ. According as the undivided act constituting vision advances more or less, the materiality of the organ is made of a more or less considerable number of mutually coördinated elements, but the order is necessarily complete and perfect. It could not be partial, because, once again, the real process which gives rise to it has no parts. That is what neither mechanism nor finalism takes into account, and it is what we also fail to consider when we wonder at the marvelous structure of an instrument such as the eye. At the bottom of our wondering is always this idea, that it would have been possible for a part only of this coördination to have been realized, that the complete realization is a kind of special favor. This favor the finalists consider as dispensed to them all at once, by the final cause; the mechanists claim to obtain it little by little, by the effect of natural selection; but both see something positive in this coördination, and consequently something fractionable in its cause—something which admits of every possible degree of achievement. In reality, the cause, though more or less intense, cannot produce its effect except in one piece, and completely finished. According as it goes further and further in the direction of vision, it gives the simple pigmentary masses of a lower organism, or the rudimentary eye of a Serpula, or the slightly differentiated eye of the Alciope, or the marvelously perfected eye of the bird; but all these organs, unequal as is their complexity, necessarily present an equal coördination. For this reason, no matter how distant two animal species may be from each other, if the progress toward vision has gone equally far in both, there is the same visual organ in each case, for the form of the organ only expresses the degree in which the exercise of the function has been obtained.

But, in speaking of a progress toward vision, are we not coming back to the old notion of finality? It would be so, undoubtedly, if this progress required the conscious or unconscious idea of an end to be attained. But it is really effected in virtue of the original impetus of life; it is implied in this movement itself, and that is just why it is found in independent lines of evolution. If now we are asked why and how it is implied therein, we reply that life is, more than anything else, a tendency to act on inert matter. The direction of this action is not predetermined; hence the unforeseeable variety of forms which life, in evolving, sows along its path. But this action always presents, to some extent, the character of contingency; it implies at least a rudiment of choice. Now a choice involves the anticipatory idea of several possible actions. Possibilities of action must therefore be marked out for the living being before the action itself. Visual perception is nothing else:[50] the visible outlines of bodies are the design of our eventual action on them. Vision will be found, therefore, in different degrees in the most diverse animals, and it will appear in the same complexity of structure wherever it has reached the same degree of intensity.

We have dwelt on these resemblances of structure in general, and on the example of the eye in particular, because we had to define our attitude toward mechanism on the one hand and finalism on the other. It remains for us to describe it more precisely in itself. This we shall now do by showing the divergent results of evolution not as presenting analogies, but as themselves mutually complementary.