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Gen. Char. Bill strong, its breadth at the base equalling or exceeding its height; upper outline of cere on a level with, or rather lower than, the base of the culmen; gonys much arched, the chord of the arch equalling about half that of the culmen. Near the tip of the upper mandible is a prominent tooth on the commissure, and near the end of the lower mandible, which is truncated, is a deep notch corresponding; the end of the upper mandible is compressed, giving the situation of the tooth an inflated appearance when viewed from above. Nostrils circular, with a conspicuous central tubercle. Orbital region bare; projecting superciliary shield conspicuous, arched, but not very prominent. Tail shorter than wing, the feathers hard and stiff. Primaries very strong, elongated, tapering rapidly toward their points; only the first or first and second with their inner webs emarginated, the cutting being angular, and near the end of the quill. Tarsus never with a single series of transverse scutellæ either in front or behind; middle toe very long.

13077, ♀. ½

Subgenera

One primary only with inner web emarginated; first to second longest; first longer than fourth.

Tarsus longer than middle toe, and feathered far below the knee; first quill shorter than third. Coloration of the sexes alike; old and young slightly different in pattern and tints. Size large … Hierofalco.

Tarsus not longer than middle toe, and scarcely feathered below the knee; first quill equal to or longer than the third. Coloration of the sexes alike; old and young very different in pattern and tints. Size, very small to large … Falco.

Two primaries with inner webs emarginated; second to third longest; first shorter than fourth.

Basal joint of toes without transverse scutellæ; tarsus about equal to middle toe.

Coloration of the sexes in adult plumage very different in tints; in the young alike, the young ♂ resembling the adult ♀. Size small … Æsalon.

Basal joint of toes with transverse scutellæ; tarsus longer than middle toe.

Coloration of the sexes very different, in pattern and tints, at all ages; old and young alike. Scutellæ of the toes and tarsus interrupted at the digito-tarsal joint; tarsus much longer than middle toe. Bill small, the cere on top less than one fourth the culmen. Size small … Tinnunculus.

Coloration of the sexes alike at all ages; old and young slightly different in pattern and tints. Scutellæ of tarsus and toes uninterrupted from “knees” to claws; tarsus but little longer than middle toe. Bill large, the cere on top about one third the culmen. Size medium; form very slender … Rhynchofalco.

Subgenus HIEROFALCO, Cuvier

Hierofalco, Cuvier, 1817. (Type, Falco gyrfalco, Linn.)

Jerafalco, Boie, 1822; Kaup, 1851. (Same type.)

Gennaia, Kaup, 1847. (Type, Falco jugger, Gray.)

Species and Races

1. F. gyrfalco. Wing, 13.00–17.00; tail, 8.50–11.50; culmen, .85–1.05; tarsus, 2.10–3.00; middle toe, 1.80–2.25.45 Ground-color varying from entirely pure white to wholly dusky, but generally bluish (in adult) or grayish-brown (in young) above, and white beneath. Adult. All the markings transverse.46 No lighter nuchal band. Young. Markings of the lower surface longitudinal, the upper parts without transverse bars (except on the tail47).

a. Lower parts with white predominating, or wholly white.

Lower tail-coverts never with markings. No tinge of blue anywhere on the plumage, the ground-color of which is entirely pure white at all ages.

1. Adult. Upper parts, excepting head and neck, with transverse crescentic bars of dark plumbeous; lower parts immaculate, or else without well-defined markings. Young. Upper parts with longitudinal stripes of dark plumbeous; lower parts usually conspicuously striped. Hab. Greenland (in the breeding-season); in winter, occasionally wandering into the northern portions of Europe and North America … var. candicans.

Lower tail-coverts always with markings. A tinge of ashy-blue more or less prevalent above. Young dusky above.

Head and neck above abruptly lighter than the back. Young plain grayish-brown above, with conspicuous whitish borders to the feathers.

2. Adult. Upper parts white, passing into bluish posteriorly; everywhere (except on head and neck) with sharply defined, transverse (not crescentic, but continuous) bars of dark plumbeous. Abdomen and flanks with transverse spots of the same. Young without irregular light mottling to the plumage above, and with broad longitudinal stripes beneath. Hab. Iceland and Southern Greenland, in the breeding-season; in winter, south into Northeastern United States, and Northern Europe … var. islandicus.

Head and neck above abruptly darker than the back. Young (of var. sacer) variegated grayish-brown above, without light borders to the feathers.

3. Adult. Top of the head streaked with whitish; back with sharply defined, continuous, narrow transverse bars, of creamy-white. Hab. Interior regions of Continental Arctic America (Slave Lake, Yukon, and McKenzie River district) … var. sacer.

4. Adult. Top of head not streaked with whitish; back without sharply defined bars of the same. Hab. Continental Arctic Europe (Scandinavia) and Siberia. Migrating south, in winter, to Bengal (Hardwicke) … var. gyrfalco.48

b. Lower parts with dusky predominating, or wholly dusky.

5. Adult. Almost entirely dusky, without well-defined markings anywhere. Hab. Littoral regions of the Hudson Bay Territory and Labrador … var. labradora.

2. F. lanarius. Wing, 11.50–16.00; tail, 6.60–9.50; culmen, .70–1.00; tarsus, 1.90–2.40; middle toe, 1.65–2.00. Ground-color varying from pale grayish-plumbeous to dark sepia-brown; beneath white, with sparse markings, these coalesced into a broken patch on the flanks. Adult. Above obscurely barred transversely with pale ashy and brownish-dusky, the former prevailing posteriorly, the latter anteriorly; a lighter nuchal band. Spots on the sides and flanks transverse. Young. Above brown, varying from grayish-drab to dark sepia, the feathers usually bordered with paler (rusty in youngest individuals); markings beneath all longitudinal.

a. Outer webs of tail-feathers with large well-defined light spots; outer webs of the primaries sometimes with light spots on the basal portion; secondaries without distinct spots on the outer webs. Lower tail-coverts immaculate.

Wing, 13.65–16.00; tail, 8.40–9.50; culmen, .85–1.00; tarsus, 1.95–2.15; middle toe, 1.85–1.95. Top of the head white, with narrow streaks of dark brown. Hab. Central and Eastern Europe, Western Asia, and adjoining portions of Africa … var. lanarius.49

b. Outer webs of tail-feathers without distinct light spots, or without any at all; outer webs of primaries with no trace of spots; secondaries with light spots on outer webs. Lower tail-coverts sparsely spotted.

Wing, 12.00–14.25; tail, 7.60–9.00; culmen, .75–.90; tarsus, 2.15–2.40; middle toe, 1.70–2.00. Top of head brown, with narrow black streaks. Adult. Above with obscure transverse spots of bluish. Young. Above with feathers bordered with rusty … var. polyagrus.

Wing, 11.50; tail, 6.60; culmen, .70; tarsus, 1.90; middle toe, 1.65. Above uniform dark brown, with a faint plumbeous cast, the feathers without trace of light or rusty edges; outer web of tail-feathers without trace of light spots. Hab. Mexico … var. mexicanus.50

Wing, 13.60–14.30; tail, 8.25–9.00; culmen, .80–.87; tarsus, 1.85–1.90; middle toe, 1.85–1.90. Colors similar to the last; entire auriculars white; mustache narrow and conspicuous. Hab. Southern Asia … var. jugger.51

The only point of difference in the external anatomy between the Lanner Falcons and Gerfalcons consists in the different degree of feathering on the upper part of the tarsus; this is much denser and extends farther down and more around the posterior face in the Gerfalcons, but they, being inhabitants of a very northern latitude, need this protection against the rigor of the climate. These slight specific differences are illustrated by the figures on page 1430. The same difference is observable in many birds whose habitat extends through a great range of latitude, as, for instance, the Pediocætes phasianellus, the northern race of which has the feathers covering the base of the toes so long as to reach beyond the claws and nearly conceal them, while in the southern form (var. columbianus) the toes are almost completely naked.

My determination of the number and character of the geographical races of F. gyrfalco is the result of a very careful critical examination of over sixty specimens, aided by the important conclusions of Mr. Hancock (Annals and Magazine of Natural History, 2d ser., XIII, 110; London, 1834), Schlegel (Falcones, Muséum d’Histoire Naturelle des Pays-Bas, 1862), Pelzeln (Uebersicht der Geier und Falken der Kaiserlichen ornithologischen Sammlung, April, 1863), and Alfred Newton (History of British Birds, revised ed., part 1, June, 1871, pp. 36–52, and Proc. Acad. Nat. Sc. Philadelphia, July, 1871, pp. 94, 95), in their important papers bearing upon this subject, which, though they each express the peculiar individual views of the writer, together clear up pretty satisfactorily the problem of the number, character, and habitats of the several races, as well as the different phases of variation to which each is subject.

43139, ♀. ½

Falco sacer.

43139, ♀. ¼

5482, ♀. ½

Falco polyagrus.

43139, ♀. ½

5482, ♀. ½

In studying the F. lanarius, I have experienced most discouraging difficulties from the want of sufficient series of the Old World races, and from the unsatisfactory character of most descriptions and figures of them, besides being much perplexed by the confusion of their synonymy by different authors. In consequence of this, my diagnoses of the four races of which alone I have seen examples may be very unsatisfactory as regards the characters by which they may be most readily distinguished. Having seen the adult of only a single one of these four races, I am therefore compelled to base my differential characters upon the immature stages.

In addition to the four races of F. lanarius characterized above, there are several geographical forms belonging to the Old World, chiefly intertropical Asia and Africa. These are the var. babylonicus, Scl. and Irby, (Gray’s Hand List, I, p. 20, No. 173,) of Southeastern Europe and Western Asia; var. barbarus, L. (Gray’s Hand List, p. 20, No. 174), of Northern Africa; and var. tanypterus, Licht. (Gray’s Hand List, No. 175), of both the preceding regions, which Mr. Gurney writes me “is simply the intertropical race of F. lanarius, from which it only differs in being of a darker shade throughout.” The F. saker, Schleg. (Gray’s Hand List, No. 176), seems, to judge from the descriptions and figures which I have seen, to be also merely a form of the same species, but I have seen no specimens of it.

Falco (Hierofalco) gyrfalco, Linn

Var. candicans, Gmelin

WHITE GERFALCON

Accipiter falco freti hudsonis, Bris. Orn. I, 356, 1763. A. gyrfalco, Briss. Orn. I, 370, pl. xxx, f. 2, 1763. Falco rusticolus, Fabr. Faun. Grœn. p. 55, 1780.—Lath. Syn. Supp. I, 15, 1781. F. candicans, Gmel. Syst. Nat. p. 275, 1788.—Daud. Tr. Orn. II, 101, 1800.—Benick, Isis, 1824, 882.—Schleg. Krit. Ubers. p. 1, 1844.—Bonap. Rev. Zool. 1850, 484; Consp. Av. p. 33.—Cassin, Proc. Ac. Nat. Sc. Phil. 1855, 278; Birds N. Am. 1858, 13.—Strickl. Orn. Syn. I, 77, 1855.—Blasius, Cab. Jour. 1862, 43 (thinks all boreal ones same in Europe and America).—Elliot, Birds N. Am. pl. xii. Hierofalco candicans, Cuv. Reg. An. ed. 1, I, 312, 1817; ed. 2, I, 323, 1829.—Less. Man. Orn. I, 80, 1828; Tr. Orn. p. 97, pl. xvi, p. 2.—Gray, Hand List I, 18, 1869. Falco islandicus, Lath. Ind. Orn. p. 32, 1790; Syn. I, 71, A, B; Gen. Hist. I, 72, A, 1821.—Steph. Zool. XIII, pt. ii, p. 39, 1826.—Gould, B. Eur. pl. xix.—Aud. Birds Am. 1831, pl. ccclxvi. F. buteo β, Lath. Ind. Orn. p. 24, 1790; Gen. Hist. I, 80, A. F. lagopus, β, Lath. Ind. Orn. p. 19, 1790; Syn. Supp. I, 36; Gen. Hist. I, 68, A. F. grœnlandicus, Daud. Tr. Orn. II, 157, 1800. Hierofalco grœnlandicus, Brehm. Voy. Deutsch, I, 16, 1831. F. gyrfalco, Bonap. List, p. 4, 1838.

Sp. Char. Adult (♀, 18,577, Greenland; Univ. Zool. Mus. Copenhagen). Ground-color entirely pure white; whole upper surface (posterior to the nape) with transverse crescentic bars of dark plumbeous-brown, generally about two on each feather, the first concealed by the feather which overlaps. Primaries crossed at regular intervals with quadrate spots of the same tint, these becoming fused toward ends of quills, forming a terminal dusky space of two or three inches in extent; tips of all the quills narrowly white; the black bars do not extend quite to the primary coverts, and decrease both in extent and regularity toward the base. Middle tail-feathers crossed with seven or eight imperfect bars of dusky, the shafts of the feathers blackish; rest of tail immaculate, the shafts pure white. Nape with a very few fine shaft-streaks of dusky. Whole lower surface of body and wing utterly immaculate. Wing-formula, 2–3–1. Wing, 16.50; tail, 9.00; culmen, 1.05; tarsus, 2.10–1.35; middle toe, 2.20; inner, 1.50; outer, 1.50; posterior, 1.00.

(No. 56,152, ♀, Greenland; Schlüter Collection.) Head above, occiput, nape, and upper half of ear-coverts, with sparse shaft-streaks of black, these most numerous on the latter region; primaries barred to the coverts. Tail entirely crossed by eleven plumbeous bars. Bars above clearer plumbeous. The snowy-white beneath is relieved by a few minute variable flecks of dusky upon the lower part of the abdomen, becoming larger as they approach the sides. Wing-formula, 2–3–1. Wing, 16.70; tail, 9.30.

Juv. transition stage? (♂ 56,047, “Hoher Norden”; Schlüter Collection). Markings above quite different from those of the two preceding; each feather has a large central longitudinal sagittate spot of dusky, leaving only the borders (of the exposed portion) white; on the primaries the dusky is almost confined to the terminal portion; the rump and upper tail-coverts have each feathers with a medial longitudinal stripe of dusky. The tail is immaculate, but the shafts of the middle feathers are dusky. The neck, breast, abdomen, and sides have numerous cuneate marks of dusky, one near the end of each feather. The lining of the wing, even, has a few narrow streaks. Wing, 14.75; tail, 9.40.

No. 56,049 (♀, Greenland, Schl. Coll.) is similar in pattern of markings, but above the dusky is more extended, forming the predominating color; the rump, etc., has broad sagittate spots instead of narrow stripes; the primaries are barred to the coverts; the tail is crossed by about ten continuous bands of dusky. Beneath the lanceolate spots or streaks cover the whole surface, except the anal region, lower tail-coverts, and throat. On the lining of the wing the streaks are less sparse than in the preceding, though they are by no means numerous. Wing, 15.75; tail, 9.50.

Falco candicans.

Juv. first plumage (♀, 56,053, Greenland; Schlüter Coll.). All the markings are longitudinal, instead of directly the reverse. The upper parts have longitudinal tear-shaped stripes, a medial one on each feather; they are sparse, however, on the wings; the rump has narrow shaft-lines of dusky. The tail and upper coverts are immaculate, but the shafts of all the feathers are nearly pure black. The bars on the primaries are found only immediately next the dusky terminal space. The streaks beneath are not very numerous, and are found only on the breast, upper part of abdomen, and on the sides; the nape and sides of the neck are, however, thickly streaked.

(No. 17,966, ♀, Moose Factory, Hudson Bay Territory.) In character of markings resembling the last, but the stripes are fainter and narrower; they are also less numerous. On the under parts they are wanting. Unfortunately, the tail of this specimen, which is the only North American one in the collection, is missing.

In all specimens the anal region and lower tail-coverts are immaculate.

Hab. Greenland, and continent of North America, north of Hudson Bay (breeding in latter region). Of irregular occurrence in winter throughout the circumpolar regions; Ural Mountains (Eversman); Behring’s Strait (Bannister).

LIST OF SPECIMENS EXAMINED

National Museum, 7; Boston Society, 2; Philadelphia Academy, 3; New York Museum, 6; collection of R. Ridgway, 1. Total, 19.

Var. islandicus, Sabine

ICELAND GERFALCON

Accipiter falco islandicus, Briss. Orn. I, 336, 1763. Falco islandicus, Sab. Linn. Trans. XII, 528, 1818.—Temm. Man. Orn. pt. x; 17, pt. iii, p. 9; Tab. Meth. p. 2, 1836.—Faber, Prod. Island. Orn. 1822, p. 2; Isis, 1827, 62.—Rich. & Swains. F. B. A. II, 27, 1831.—Hoy, Mag. Nat. Hist. Ser. 1, VI, 107.—Hancock, Ann. Nat. Hist. II, 247; Rev. Zoöl. 1839, 123.—Bonap. Consp. Av. p. 24.—Strickl. Orn. Syn. I, 77, 1855.—Cassin, Birds N. Am. 1858, 13. Hierofalco islandicus, Gray, Gen. B. p. 3 (ed. 2, p. 4), 1844; Hand List, I, 18, 1869. Falco candicans islandicus, Schleg. Krit. übers, p. 1, 1844. Falco lanarius, Faber, Isis, 1827, 68. Falco gyrfalco, Keyserling & Blasius, Wirbelth. Eur. p. 135, 1840.

Sp. Char. Adult (♂, Iceland; No. 12, Coll. Geo. N. Lawrence). Ground-color of the plumage dull white, gradually becoming somewhat bluish posteriorly, this color especially noticeable on the tail. Whole upper parts crossed with broad transverse bands of dark plumbeous, these bands continuous, and more than twice as wide as the pale ones, except on the upper tail-coverts and tail, where the bands of the two colors are more regularly defined and about equal; in addition to the transverse bands, the feathers anteriorly have narrow borders of white. Tail with the dark bands twelve in number; the terminal pale band is purer white than the others. The dusky plumbeous prevails on the primaries, and is unvariegated beyond the middle portion; the anterior half, however, is marked with quadrate ragged spots, of a slightly yellowish-white; all are margined terminally with purer white. Each feather of the head and neck with a narrow medial streak of dusky, but the general aspect abruptly lighter than the back; the streaks are more condensed along the upper and terminal portion of the ear-coverts. Jugulum and breast with a medial narrow streak on each feather; abdomen with more elliptical streaks; sides with circular and cordate spots, and flanks and tibiæ with transverse spots; lower tail-coverts with narrow shaft-streaks of dusky. Lining of the wing with sparse narrow streaks of dusky; under surface of primaries with white prevailing, this, however, crossed by narrow bars of dusky, these numbering about sixteen on the longest. Wing-formula, 2–3–1. Wing, 14.60; tail, 7.80; culmen, 1.00; tarsus, 2.30; middle toe, 2.00.

Juv. (No. 20,344, Iceland). Ground-color of head, neck, and lower parts, white. Upper surface grayish umber-brown, becoming paler and more grayish on the tail; each feather above sharply bordered (both webs, all round) with dull white, producing a somewhat squamate appearance; in places, a few obsolete hidden spots of yellowish-white. Tail ashy-drab (feathers somewhat paler along edges), crossed with about eleven transverse series of spots of ochraceous or creamy white; these very obsolete on middle feathers, and sharply defined only on inner webs; the last is terminal. Primaries plain brown, somewhat darker than the back, and becoming insensibly darker terminally; skirted with white, and somewhat mottled or irregularly spotted toward their bases with yellowish-white. Head and neck, each feather, with a medial streak of dusky, but white the prevailing aspect; these streaks condensed and somewhat suffused along upper border of ear-coverts, and from the lores along cheeks, forming an obsolete “mustache”; every feather beneath (including lining of wings) with a medial broad stripe of clear plumbeous vandyke-brown, the shaft pure black; under surface of primaries with transverse spaces of white, these numbering thirteen on the longest. Wing-formula, 2–3, 1. Wing, 15.00; tail, 9.20.

Hab. Iceland and Southern Greenland. Northeastern North America in winter, straggling accidentally south to the New England States; Rhode Island (Museum, Cambridge); Norway, Maine “not uncommon” (Verrill); Massachusetts (Peabody & Jillson); Long Island (Cab., G. N. Lawrence).

Falco islandicus.

No. 56,050, Greenland (Schlüter Collection), is moulting, and assuming the adult dress; the adult and young stages above described being nearly equally combined. No. 56,055, from Greenland, differs from the other young individuals which I have seen in being considerably darker. The feathers of the upper surface are not bordered with whitish, but are merely paler on their edges, along which are specks of yellowish. On the head and neck the dark streaks predominate, while the stripes below are very broad. It approaches quite nearly toward the young of var. sacer.

The only specimen of this race which I have seen from Continental North America, is a young individual, obtained during the winter of 1864–65, near Providence, R. I., taken by Mr. Newton Dexter, and now in the Cambridge Museum, where I had the pleasure of seeing it.

LIST OF SPECIMENS EXAMINED

National Museum, 5; Boston Society, 3; Philadelphia Academy, 9; Coll. G. N. Lawrence, 2; Museum Comp. Zoöl., 1; New York Museum, 5. Total, 25.

Var. sacer, Forster

MacFARLANE’S GERFALCON

Falco sacer, Forster, Phil. Trans. LXII, 1772, 383 and 423.—Coues, Birds of New England, 1868, 6.—Baird, Trans. Chicago Acad. Sc. I, ii, 271. ? Falco cinereus, Gmel. Syst. Nat. p. 267, 1789.

Sp. Char. Adult (♂, 51,689, Yukon, mouth of Porcupine River; Strachan Jones). Whole upper surface with numerous transverse bands of brownish-plumbeous and ashy-white. Anteriorly the light bars are about half the width of the dark ones; posteriorly they gradually increase, the bands of the two colors being about of equal width on the upper tail-coverts and tail; with the increase of the lighter bars, they become more ashy, and, correspondingly, the darker ones are more plumbeous; on the rump there is but little contrast between the bands of the two, causing a prevalent bluish cast. The bands are everywhere continuous, the light ones being interrupted only by the black shaft; there are generally on the anterior portions about three light bars on each feather, the last always terminal. Tail tipped with white, and crossed with equal continuous bands of hoary-plumbeous and ashy-white; the latter eleven in number, and finely sprinkled with deeper ash. Primaries brownish-plumbeous, plain past the middle portion, but on the anterior half with quadrate spots of creamy white on the outer web. Head above brownish-plumbeous, this prevailing; but along the median line the feathers are edged with buffy white; forehead dull white, this continuing back in a streaked superciliary stripe to the occiput; cheeks very thinly marked with fine streaks of dusky, this prevailing along the upper border of the ear-coverts; a deeper dusky suffusion beneath the anterior angle of the eye. Lower surface pure white; chin and throat, only, immaculate; jugulum with very sparse, narrow longitudinal streaks of blackish; sides with scattered cordate or nearly circular spots, these larger and transverse on the flanks and tibiæ; abdomen with scattered minute elliptical spots; lower tail-coverts with minute irregular sagittate or transverse spots of dusky. Under surface of the wing white; each feather of the lining with a medial tear-shaped streak of dusky; primaries crossed with narrow bars of dusky, fifteen in number on the longest. Wing-formula, 2–3–4–1–5. Wing, 13.50; tail, 8.60; culmen, .90; tarsus, 2.15; middle toe, 1.87.

♀ (43,139, Fort Anderson, May 24, 1864, “♀ and two eggs”; R. MacFarlane). Generally similar to the male. Head above conspicuously streaked, but the dusky prevailing. Above the transverse bands are less regular and continuous, anteriorly the plumbeous largely prevailing; posterior portions, however, as in the male, but on the rump the bands are more distinct. Beneath, the markings are more numerous, larger, and broader; those on the jugulum linear; those of the abdomen medially elliptical; laterally they are transversely cordate, and on the flanks in form of broad transverse spots, or broad bars; on the tibiæ and lower tail-coverts they form regular transverse bars,—on the latter, quite distant. Wing-formula, 2–3–4, 1. Wing, 15.50; tail, 9.50; tarsus, 2.15 and .80; middle toe, 1.95.

Juv. (♂, 55,400, Alaska, Nulato, February 10, 1868; W. H. Dall). Above plumbeous-umber, precisely as in young of islandicus, but on the rump having a decided ashy cast. No white edges to the feathers, as in islandicus, but, instead, numerous irregular transverse spots or obsolete ragged bars of cream-color or pale ochraceous-buff; the whole upper surface is quite thickly variegated with these irregular markings. Tail crossed with thirteen narrow bands of creamy-white, these so thickly mottled with dusky on the outer webs as to be obscure, but on inner webs they are regular and sharply defined; the last is terminal. Primaries plain dusky, skirted obscurely with paler, and marked toward bases with obsolete mottled spots of cream-color. Head streaked with dusky and creamy-white, the former predominating on upper surface, along upper edge of ear-coverts, and across the cheeks, on the latter forming a mustache; the white prevails over the ear-coverts in a broad supra-oral stripe, and on the forehead and lores. Beneath, soft dull white; chin and upper part of throat, only, immaculate; each feather with a broad medial stripe of clear dark plumbeous-brown, on the flanks and tibiæ prevailing, the whitish assuming the form of roundish spots; lining of the wing similarly marked; prevailing aspect of under surface of primaries white, crossed with narrow bars of ashy, fifteen in number on the longest. Wing-formula, 2, 3–1=4. Wing, 14.00; tail, 8.40.

Hab. Interior regions of Arctic America; Anderson River, McKenzie, Yukon, and Severn River regions. Breeding abundantly in the former district, whence numerous specimens of skins and eggs have been received by the Smithsonian Institution.

In the young specimen described, there are one or two new feathers appearing on the rump and upper tail-coverts, precisely as in the blue plumage, and proving conclusively their relationship. The species is as different from the Iceland bird in the young stage as in the mature. The most readily apparent differences are, lack of sharp white edges of feathers above, and in their stead numerous ragged transverse spots of yellowish; dark aspect of head above, etc.

Specimens vary considerably in the shades of color and distribution of the markings, but the types of the above descriptions are the lightest of the series. The darkest example is No. 43,144½ (“♀ and eggs”), Fort Anderson, May 22, 1864. In this the whole head and neck (except underneath) are continuous blackish-plumbeous, only the middle of the auriculars being faintly streaked; the back is nearly plain dusky, and even on the wings the bars are very obscure and much reduced in width. The rump is plain ashy-blue, the darker bars being nearly obsolete. The longitudinal markings on the pectoral region are enlarged into conspicuous stripes, while on the sides and flanks the transverse bars form heavy spots. The transverse bars on the tibiæ are ashy-blue; those on the crissum clear plumbeous, and regularly transverse. Wing, 15.75; tail, 9.30. Upon comparing this specimen with the figures of a pair of var. gyrfalco, by Wolf, in Newton’s Oötheca Wolleyana, I can discover no difference at all; thus it would seem that our bird occasionally closely approaches in tints and markings this race of Continental Europe, of which I have seen only one immature example, and no adults.

I cannot agree with Mr. Newton in considering the Gerfalcons of the interior of Arctic America as identical with the Iceland form, though that distinguished ornithologist considers them so in his paper in the Proceedings of the Philadelphia Academy for July, 1871, basing his conclusion upon the specimens from which the above descriptions were taken, which had been sent over to England for comparison. I have never yet seen a specimen of islandicus which could not be distinguished, by the characters given in my synopsis, from these examples, while they can be separated from that race by the characters which Mr. Newton himself gives, in his diagnostic table in the paper above cited, for distinguishing the adults of islandicus and gyrfalco.

The var. sacer is evidently separable from both islandicus and gyrfalco, and about as much related to one as to the other; combining the size and proportions of the former with the colors of the latter, while in the wide amount of individual variation of plumage its lighter extreme approaches one, while its darkest phase approximates as closely to the average plumage of the other.

LIST OF SPECIMENS EXAMINED

National Museum, 6.

Var. labradora, Audubon

BLACK GERFALCON

Falco labradora, Aud. B. Am. pl. cxcvi, 1831.

Sp. Char. Adult (♀ breeding plumage? 30,375, Rigolet, Labrador; Mr. Conolly). Ground-color of the plumage uniform, very deep, clear, dark plumbeous-brown, continuously uniform above; larger scapulars, secondaries, secondary coverts, and primaries more dilute along edges, however, the tint palest and broadest terminally. Tail perfectly uniform, except at the end; the tip being narrowly whitish, and about half an inch anterior to this, a transverse series of hidden irregular transverse creamy-white spots. The head (except beneath) is unvariegated. Beneath, the dark tint inclines more to blackish clove-brown, more dilute on the tibiæ; feathers edged laterally with white, this prevailing on the throat, but everywhere else far less than the dusky in amount; on the tibiæ and lower tail-coverts the white is in the form of irregular spots. Anal region unvariegated; lining of the wing with circular spots of white along the outer webs of the feathers. Under surface of primaries with plumbeous prevalent, but this crossed with mottlings of whitish, forming transverse bars; but terminally and basally they become confused or lost. Wing-formula, 2, 3–1, 4. Wing, 16.20; tail, 9.50; tarsus, 2.00–.90; middle toe, 2.05; inner, 1.50; outer, 1.50; posterior, .90.

Hab. Labrador; south and westward in winter, and shores of Hudson Bay.

Nos. 17,063 (♀, Quebec, W. Cooper) and 34,960 (♀, Fort Nescopec, Labrador) differ from the preceding in having ten small narrow transverse spots of reddish-white on the tail-feathers, forming as many indistinct bands; these spots touch neither the shaft nor the edge of the feather, and are almost concealed, unless the tail is spread; on the latter specimen they are very obsolete, the subterminal one only being distinct, as in the specimen selected for description. The upper tail-coverts also show faintly indicated spots, and the former specimen has the wing-coverts with very narrow irregular spots on the edge of the feathers. In this specimen there is also one feather in the scapulars which has broader white edges; it also has the white below about equal to the black in amount; the anal region, however, in all, is unvaried blackish, and the transverse oblique bands on the lower tail-coverts are a constant feature.

No. 41,185 (♀, Fort Nescopec, Labrador; H. Conolly) is the darkest of all. In this the blackish plumbeous-brown is uniform over the whole surface; even the throat is unvariegated. Abdomen with a few of the feathers edged with white, and sides with a few small circular spots of the same; lower tail-coverts transversely spotted with white; tibiæ scarcely variegated, showing only narrow indistinct whitish edges. Mottling on inner webs of primaries reduced so as to be scarcely visible. Tail with the usual number (two) of irregular whitish bars,—one terminal, the other near the end.

LIST OF SPECIMENS EXAMINED

National Museum, 2; Boston Society, 1. Total, 3.

Habits. In treating of the general habits of the Gerfalcons of North America it will not be necessary, nor will it be possible, to give the distinctive peculiarities belonging to the several forms in which these Falcons occur. Whether, on account of their variations of plumage, we consider them as races or as specifically distinct, does not affect their history in this respect. There is no good reason for presuming that they have any very noticeable variations as to any of their habits, although certain writers claim for some of them certain well-marked peculiarities of character.

In the matter of geographical distribution they are all, for the most part, rarely seen, even in midwinter, south of the 50th parallel of north latitude, and are found in the summer as far north as the Arctic Ocean. The Gerfalcon of the McKenzie River region, occurring from the Slave Lake to Anderson River and the Yukon, is the form elsewhere given as the F. sacer. Along our eastern coast region occurs another form, the F. labradora, which is the bird met with in Labrador, and described by Mr. Audubon. The F. candicans or grœnlandicus is a form peculiar to Greenland, visiting also, in the winter, the Hudson’s Bay region; while the F. islandicus, a well-known European form, occurs in Greenland also, and occasionally farther south.

Holböll, in his account of the birds of Greenland (Isis, 1845), appears to recognize but one species of Gerfalcon as occurring there, to which he gives the name of islandicus. This is, he states, the most abundant Falcon in Greenland, and is equally common in the northern and in the southern parts. Their great variations in color he regarded as indicative of differences in ages to only a very limited extent, and as in no respect specific. These differences in color were found among both nestlings and breeding birds, white and dark birds being found together in both circumstances. The white birds were more numerous in Northern Greenland, and the dark ones oftener seen in the southern portion.

He found the young birds moulting throughout the winter. On the 4th of January, 1840, he shot a young female that showed signs of moulting about the head and neck, with a striped white appearance from the sprouting feathers. The ovaries were quite well developed, and it was evident that the birds of this species breed in the first season after their birth. Holböll adds that they breed in January, that their eggs are of nearly the same color as those of the Ptarmigan, but are twice as large. They nest usually in inaccessible cliffs. They prey chiefly upon water-fowl and Ptarmigans, and usually build near “bird rocks,” from which they obtain the young without much trouble. He mentions having once seen one with a young Larus tridactylus in each foot, and another with two Tringa maritima carried in the same manner. Its rapidity of flight Holböll did not regard as very great. He had for years kept pigeons, and only lost two young birds, which were seized when at rest. Almost every day, especially in October and November, these Falcons would chase the old Pigeons unsuccessfully, and were often shot when they followed them too near the house. They were not particularly shy, and were occasionally decoyed and killed by throwing a dead bird towards them.

During the summer they are most numerous along the bays, especially where there are “bird-rocks” near. In September they go southerly along the coast, and also in October and November. At this time they are not rare, and approach the houses of the Danes, near which they are often seen fighting with the Ravens. Their spring migrations are not so regular as they are in the autumn, or perhaps at this time they do not approach the houses so frequently. When they are near the settlements, it is noticed that in the morning they fly towards the south, and in the evening towards the north.

Richardson speaks of the Gerfalcon as a constant resident in the Hudson Bay territory, where it is known as the Speckled Partridge-Hawk, and also as the Winterer. Its southern limit he could not give, but he never met with it south of 52°. He traced it northward to the coast of the Arctic Sea, and probably to the most northern Georgian islands. He cites Captain Sabine as authority for its occurring as far north as latitude 74° on the west coast of Greenland. Richardson often met with it during his journeys over the Barren Grounds, where its habitual prey was the Ptarmigan, and where it also destroyed Plover, Ducks, and Geese. He relates that in the middle of June, 1821, a pair of these birds attacked him as he was climbing to the vicinity of their nest, which was built on a lofty precipice on the borders of Point Lake, in latitude 65° 30′. The bird flew in circles, uttering loud and harsh screams, stooping alternately with such velocity that their motions through the air produced a loud rushing noise. They struck their claws within an inch or two of his head. Keeping the barrel of his gun close to his cheek, and suddenly elevating its muzzle when they were in the act of striking, he found that they invariably rose above the obstacle with the rapidity of thought, showing equal power of motion. They bore considerable resemblance to the Snowy Owl, but their flight was much more rapid.

Mr. MacFarlane, in the memoranda of his collections in the neighborhood of Anderson River and Fort Anderson, furnishes notes of eighteen nests of the Gerfalcon obtained by him in that region. With only two exceptions, these were placed near the tops of pines, or other trees, at distances from the ground varying from ten to twenty-five feet. In some instances the nest was placed on the very top of the tree, in others on a lower limb against the trunk. They were composed of twigs and small branches, and lined with mosses, hay, deer’s hair, feathers, and other substances. The parents were always very much excited whenever their nests were approached, making a great noise, and not unfrequently their loud screams drew attention to nests that would otherwise have escaped notice. In one instance a nest had been built on a ledge of rocks thirty miles northwest of Fort Anderson. It was composed of a few withered twigs, and lined with mosses and hay. It was found on the 27th of May, and contained two eggs nearly fresh, and two in a state of greater development. One nest, placed on a broad branch of a tree, near the trunk, was of considerable size. Another nest was on the ground, on the side of a steep and high hill. The earliest date of finding these nests is given as the 10th of May. The eggs then found were fresh. The ground at that time was still thickly covered with snow, and the weather was very cold. In a nest found five days later the eggs contained partially developed embryos. In nearly every instance the eggs seem to have been in different stages of development in the same nest. In some, young birds were in the same nest with eggs only partially developed, and in another an egg perfectly fresh was in the same nest with others nearly ready to hatch. A nest found July 3 contained young about two days old; another, on May 27, had eggs with large embryos; and one, on June 25, had young nearly ready to fly.

Mr. Donald Gunn claims that this Falcon is the only Hawk that is resident in the Arctic regions throughout the year. It is known to the Indians by the name of Pepunesu, and this name is applied to it because it passes the winter with them. It is a very powerful bird, and commits great havoc among the Partridges, so much so that in former times the Hudson Bay Company gave a reward of a quart of rum to every hunter who brought in the head of one of these Falcons. All the other Hawks are only summer visitors.

Mr. Bannister was informed by the residents of St. Michaels that a Hawk, presumed to be this species, is not unfrequent there, though he did not happen to meet with it. On his voyage home, on the 21st of October, 1866, when off the coast of Kamtschatka, north of Behring’s Island, one alighted in the rigging of the ship, and continued with them for several hours.

Although very rare in any part of the United States, occasional individuals have been taken in different localities, and in one instance a pair was known to breed for several successive seasons in Vermont. This information I have from Mr. Clarence King, who, when a lad at school in the town of Dummerston, observed a pair nesting among some high cliffs, and informed me of the fact at the time of the occurrence. One of these birds is recorded by Mr. Lawrence as having been taken on Long Island in the winter of 1856.

Mr. Boardman gives it as occurring near Calais in winter, but very rare. Professor Verrill found them not uncommon in Oxford County, Me., where they were frequently seen during winter, flying about the extensive meadows near Norway; but they were very shy and watchful, and it was hardly possible to procure a specimen. It is very unusual in Eastern Massachusetts, and only very rarely and occasionally have specimens been taken. Mr. Jillson obtained a specimen, in 1840, at Seekonk. One was shot, in 1864, near Providence, R. I., by Mr. Newton Dexter.

Mr. Audubon relates that, August 6, 1833, his son, John W. Audubon, found a nest of this Falcon among some rocky cliffs near Bras d’Or, Labrador, containing four young birds ready to fly, two of which were procured. The nest was placed among the rocks, about fifty feet from their summit and more than a hundred from their base. It was inaccessible, but, having been examined from above, was seen to be empty. It was composed of sticks, sea-weeds, and mosses, was about two feet in diameter, and was almost flat. Its edges were strewed with the remains of their food, and beneath the nest was an accumulation of the wings of Ptarmigans, Mormons, Uriæ, etc., mingled with large pellets of fur, bones, and various substances.

Their flight is spoken of as similar to that of the Peregrine Falcon, but more elevated, majestic, and rapid. Their cries were also like those of that Falcon, being very loud, shrill, and piercing. Occasionally this bird was seen to alight on one of the high stakes placed on the shore. There it would stand, in the position of a Tern, for a few moments, and then would pounce upon a Puffin, as the latter bird was standing at the entrance of its burrow, unaware of the approach of its enemy. The weight of the Puffin seemed to form no impediment to the Hawk in its flight.

The European Gerfalcons are said to seldom appear south of the 52d parallel of latitude, or north of 74°. They are nowhere numerous, and were formerly much sought for, and purchased, at immense prices, for purposes of falconry. Great differences were supposed to exist in regard to the habits and other peculiarities of the several races. The Iceland Falcons commanded the highest prices, and were regarded as a species quite distinct from the F. gyrfalco. The former was much the more valuable, both as more rare, and as a bird of higher courage and of a more rapid and bolder flight, and a bird that could, on that account, be “flown” successfully at larger game.

The Gerfalcons, in Europe, build on the rocky coasts of Norway and Iceland, and are said to defend their young with great courage and determination. They are comparatively rare in the British Islands, especially the more southern portions. Even in the Orkneys it is only an occasional visitor.

All the eggs of the several forms of Gerfalcon that I have seen present common characteristics, and do not differ from each other more than eggs known to belong to the same species of Hawk are frequently found to vary. One from Greenland, presumed to belong to the candicans, measures 2.37 inches in length by 1.71 in breadth. The predominant color of its markings is a deep reddish-brown, very generally and nearly equally diffused over its surface, concealing the ground-color, which is lighter and of a yellowish-brown shade.

An egg of the islandicus, from Iceland, has the same measurements, but is so slightly yet uniformly marked with light yellowish-brown as to seem to be of one color only,—a light brown, shaded with yellow.

An egg from Norway, of the form gyrfalco, is 2.42 inches in length, 1.71 in breadth, has a ground-color of a dirty yellowish-white, and is marked with spots, dottings, and confluent blotches of yellowish-brown, more so about the larger end.

The series of eggs of Falco sacer in the Smithsonian Collection exhibits the following range of variation in size, color, and markings: length, from 2.30 to 2.45 inches; breadth, 1.60 to 1.90 inches; ground-color usually a light reddish-ochre, varying to pinkish on the one hand, and to rufous on the other. They are usually sprinkled all over with small spots, which are sometimes not distinguishable from the ground-color when this is very deep, and again larger and quite conspicuous.

An egg of the variety candicans, from Greenland (No. 2,606, S. I.), measures 2.25 inches by 1.80. In color and in markings it is like the average eggs of variety sacer, namely, pale rufous, sprinkled over with a slightly deeper shade.

Falco lanarius

Var. polyagrus, Cassin

AMERICAN LANNER; PRAIRIE FALCON

Falco polyagrus, Cassin, B. Cal. & Tex. 1853, 88.—Ib. P. A. N. S. 1855, 277; B. N. Am. 1858, 12.—Heerm. Pacific R. Rep’t, II, 1855, 31.—Kennerly, P. R. R. III, 1856, 19.—Coop. & Suckl. P. R. R. XII, 1860, 143.—Coues, P. A. N. S. 1866, 7.—Strickl. Orn. Syn. I, 1855, 85.—Dresser, Ibis, 1865, 323.—Gray, Hand List, I, 1869, 20. Falco lanarius, var. mexicanus, Ridgway in Coues’ Key, 1872.

Sp. Char. Adult (♂, No. 59,063, Wahsatch Mountains, Utah, May 23, 1868; parent of eggs; L. E. Ricksecker). Above cinereous-drab, becoming gradually paler and more bluish posteriorly, barred, indistinctly, everywhere with a more dusky tint, the shafts of all the feathers blackish; anteriorly the darker shade predominates, while posteriorly the bluish prevails; on the anterior portions the light bars are much restricted in width, and of a more ochraceous tint. Tail plain, very pale ashy-drab, narrowly tipped with reddish-white, this changing to pale rusty on the middle pair; the concealed portion of the feathers outside the shaft show obsolete, or faint traces of, darker bars, which on the middle pair are apparently about eleven in number. On the inner webs the paler bars become broader than the darker ones, and incline to ochraceous in tint, the lateral feather being edged externally with this color. Primaries plain ashy-drab, with a hoary tinge, growing insensibly darker terminally, and with a slightly paler apical margin. Head and neck above, dark umber-brown, with conspicuous shaft-streaks of black. Lores, a broad superciliary stripe (somewhat interrupted above the eyes), white, finely and sparsely streaked, the two stripes confluent across the occiput; a broad heavy “mustache” from the lores and rictus downward and obliquely backwards, across the maxilla, and a wider postocular stripe, like the crown. Beneath continuous white, with a faint ochraceous tinge on the abdomen and crissum; abdomen and sides of the breast with a few scattered, small, ovate spots of vandyke-brown; sides transversely spotted with vandyke-brown, the spots coalesced into a broken patch on the flanks; outside of the tibiæ with transverse spots of the same. Axillars plain, clear vandyke-brown, with a few nearly obsolete rusty specks near their ends; lining of the wing clear white, the feathers with central spaces of dusky-brown, which toward the edge become aggregated into a longitudinal patch; inner webs of the primaries with broad transverse spots of white, which reach nearly to the shaft; they are about thirteen in number on the longest quill. Feet yellow; base of the bill tinged with the same. Wing-formula, 2, 3–1, 4. Wing, 12.00; tail, 7.50; tarsus, 1.90; middle toe, 1.70; outer, 1.22; inner, 1.12; posterior, .77.

♀ (not adult? 18,258, Fort Buchanan, New Mexico; Dr. Irwin). Above continuous umber-drab, growing gradually lighter posteriorly, the tail being pale drab; no transverse bars (except a few concealed obsolete ones on back and secondaries), but all the feathers faintly bordered with paler rusty-brown, these edgings being on upper tail-coverts almost white. Tail tipped with creamy-white, and with many transverse spots or broad bars of the same on inner webs, outer feather irregularly skirted with the same, and all decidedly paler than the ground-color along their edges. Head as in the male, but forehead white, and superciliary stripe more continuous. Breast and abdomen with longitudinal lanceolate or cuneate streaks of dark vandyke-brown; patch of same on flanks more continuous than in the male; axillars unvariegated clear dark vandyke-brown; longest primary with eleven transverse spots of white; posterior outer face of tibiæ with sagittate spots of dark brown. Wing-formula, 2, 3–1, 4. Wing, 14.25; tail, 8.00; tarsus, 2.10; middle toe, 2.00.

Juv. (♂, 32,207, South Fork of the Platte River, July 19, 1838; C. S. McCarthy). Above darker umber than the last, each feather distinctly bordered terminally with rusty-ochraceous. Beneath with a deeper cream-colored tinge, streaks blacker; flank-patch more conspicuous and uniform; axillars unvariegated dusky. Wing-formula, 2, 3–1=4. Wing, 13.25; tail, 7.25.

Hab. Western division of North America, eastward to Illinois; Oregon to Lower California, and Texas. Localities: Texas, San Antonio and Eagle Pass (Dresser); Arizona (Coues).

The different stages of plumage are in this by no means so well defined as in other species, there being nearly the same general appearance in all. There is, also, very little variation in different specimens of the same age. No. 8,504, (♀, Dalles, Oregon; Dr. George Suckley) has the black markings on the sides of the breast more circular, and the vandyke-black of the axillars with a few circular white spots on the edges of the feathers. Wing, 14.50; tail, 8.40. Nos. 17,204 (♀, San José, Lower California; John Xantus, January, 1860) and 18,258 (♂ ? Fort Buchanan, N. M.) have the upper surface almost perfectly continuous grayish-drab, the first absolutely unvariegated by markings, though the feathers fade a little on edges. Beneath, the white is very pure; the streaks are numerous, sharply defined and longitudinal. Wing, 13.25; tail, 7.50 (17,204).

The American Lanner Falcon is so very closely related to the Lanners of Europe and Asia (var. lanarius and var. jugger) that it is very difficult to indicate the differences which separate them. The two Old World forms above named are more unlike each other than they are from the two American races; the var. jugger differing from mexicanus apparently only in larger size; and the var. lanarius, more like polyagrus than it is like either jugger or mexicanus, differs from polyagrus mainly in the greater amount of white on the plumage, this imparting a lighter aspect to the pileum, and causing a greater development of the light spots on the outer webs of the primaries and rectrices.

Falco polyagrus.

The var. polyagrus, compared with var. lanarius, is much darker, having, at all ages, the crown uniformly brown, with darker streaks, instead of having these streaks upon a white ground. The “mustache” is more distinct in the American bird, while in the European the bands on the tail are much more distinct, and the spots forming them are on the outer webs, as well as on the inner, instead of on the latter alone; the dark bars between the light spots are in the American bird much narrower and more numerous, and in the young the light ones come to the edge of the web, instead of being enclosed within the dark color. Two very young birds (i.e. in first perfect plumage) appear almost identical until closely examined, the chief differences being a lighter tint to the crown in the European, and heavier dark stripes on the breast, besides the peculiar character of the tail-spots, which are always distinctive. In shades of color, there is not the slightest difference.

I have seen no specimen of any of the Old World forms in the plumage corresponding to that transversely barred above, described here as the adult, though figures of the adult lanarius indicate a very similar plumage. The series of the latter race at my command is unfortunately limited to a very few immature specimens. One marked “ad.” (56,051, Hungary; Schlüter Coll.) measures as follows: Wing, 14.50; tail, 8.00; culmen, .83; tarsus, 1.90; middle toe, 1.80. Its colors are as described in the synopsis (p. 1429) for the young bird.

The var. mexicanus and var. jugger, which are both much darker, and more uniform in the coloring of the upper parts, than var. polyagrus, are more nearly alike; in fact, the only tangible difference that I can find between a specimen of the former in the Museum of the Boston Society of Natural History (No. 1,438, ♂, Juv. Lafr. Collection; “Mexico”) and two examples of the latter in the New York Museum, consist in the larger size of the var. jugger (see synopsis), besides its whiter cheeks and more isolated and distinct “mustache.” A direct comparison of these two races may show other tangible points of distinction, or, on the contrary, may show even these slight distinguishing features to be inconstant. The former result is, however, most reasonably to be expected.

LIST OF SPECIMENS EXAMINED

National Museum, 9; Boston Society, 2; Philadelphia Academy, 4; Museum Comp. Zoöl. 1; G. N. Lawrence, 2; R. Ridgway, 5. Total, 23.

Habits. This is an exclusively western species, occurring from the valley of the Mississippi to the Pacific coast. Specimens have been obtained as far east as Illinois. Several others have been taken on the Upper Missouri and the Yellowstone Rivers, in Nebraska, at Fort Thorne, New Mexico, and on the Little Colorado River. A specimen was shot by Dr. Heermann on the Farallones, on the California coast; but Dr. Cooper thinks it rarely visits the coast border, though he several times saw, near San Diego, a bird which he supposed to belong to this species. At Martinez, in December, 1863, he succeeded in shooting one as it flew from its perch at the approach of the wagon in which he was riding.

It is said to extend its migrations in summer to the Upper Columbia, avoiding the densely forest-clad regions. Dr. Heermann saw a young unfledged individual at San Francisco, from which it may be inferred that a few may breed within the State.

The first individual of this species was taken by Dr. Townsend during his trip across the continent, in 1834. It was obtained among the mountainous regions of Oregon, near the sources of the Platte River. Mr. Cassin states that Dr. Heermann procured several specimens in the Sacramento Valley.

Mr. Cassin remarks that this species, except in its greatly superior size and strength, bears a very close resemblance to the well-known Jugger Falcon of India, a bird much used for the purposes of falconry.

Dr. Kennerly, who procured a single specimen of this species while his party was encamped on the Little Colorado, found it busily engaged in seeking its prey among the bushes that grew along the river-bank. It was shy, and was procured with difficulty.

Dr. Suckley speaks of this Hawk as not at all rare in Oregon. He procured a specimen of it at Fort Dalles, in the beginning of the winter of 1854–55, which had been killed in the act of carrying off a barn-yard fowl of about its own weight, and which it had just seized near the door of a dwelling-house,—an act demonstrative of a union of courage, ferocity, and strength inferior to none of its congeners.

Dr. Cooper characterizes this as one of the shyest of Hawks, as it is also one of the swiftest, flying with rapid flappings of the wings. It seems to prefer the borders of prairies, where it catches hares, quails, and even larger game.

Mr. Ridgway informs me that this Hawk was seen by him in Southern Illinois, near Mt. Carmel, September 27, 1871. It had been obtained once before within the limits of Illinois, but in the northwestern part of the State, at Rock Island, by I. Dickenson Sergeant, of Philadelphia, and presented by him to the Academy of Natural Science.

Its nest and eggs were taken in Utah by Mr. Ricksecker. I have no notes in regard to the former. A finely marked specimen of one of the eggs procured by him is in my cabinet. It measures 2.15 inches in length by 1.65 in breadth. It is of a somewhat less rounded-oval shape than are the eggs of the anatum. The ground-color is a rich cream, with a slightly pinkish tinge, and is beautifully marked with blotches of various sizes, shapes, and shades of a red-brown tinged with chestnut, and with occasional shadings of purplish. These are confluent about one end, which in the specimen before me chances to be the smaller one. It very closely resembles the eggs of the European F. lanarius.

An egg in the Smithsonian Collection (15,596), taken at Gilmer, Wyoming Territory, May 13, 1870, by Mr. H. R. Durkee, has a ground-color of pinkish-white, varying in two eggs to diluted vinaceous, thickly spotted and minutely freckled with a single shade of a purplish-rufous. In shape they are nearly elliptical, the smaller end being scarcely more pointed than the larger. They measure 2.27 by 1.60 to 1.65 inches. The nest was built on the edge of a cliff. Its eggs were also taken by Dr. Hayden while with Captain Raynolds, at Gros Vent Fork, June 8, 1860.

Subgenus FALCO, Mœhring

Falco, Mœhring, 1752. (Type, Falco peregrinus, Gm. = F. communis, Gm.)

Rhynchodon, Nitzsch, 1840. (In part only.)

Euhierax, Webb. & Berth., 1844. (Type, Falco—?)

Icthierax, Kaup, 1844. (Type, Falco frontalis, Daud.)

51293, ♂. ¼

F. aurantius.

52814, ♀.

F. rufigularis (nat. size).

51293, ♂. ½

F. aurantius.

51293, ♂. nat. size.

F. aurantius.

52814, ♀.

F. rufigularis (nat. size).

The following synopsis of the three American species of this subgenus may serve to distinguish them from each other, though only two of them (F. aurantius and F. rufigularis) are very closely related. The comparative characters of the several geographical races of the other one (F. communis), which is cosmopolitan in its habitat, being included under the head of that species, may explain the reasons why they are separated from each other.

Species and Races

A. First and second quills equal and longest; first with inner web emarginated, second with inner web slightly sinuated. Young with longitudinal stripes on the lower parts. Adult and young stages very different.

1. F. communis. Wing, 11.50–14.30; tail, 7.00–8.50; culmen, .72–.95; tarsus, 1.65–2.20; middle toe, 1.80–2.30.52 Second quill longest; first shorter than, equal to, or longer than third. Adult. Above plumbeous, darker anteriorly, lighter and more bluish posteriorly; anteriorly plain, posteriorly with darker transverse bars, these growing more sharply defined towards the tail. Beneath ochraceous-white, varying in tint from nearly pure white to deep ochraceous, those portions posterior to the jugulum transversely barred, more or less, with blackish or dark plumbeous; anterior lower parts (from the breast forward) without transverse bars. Young. No transverse bars on the body, above or below. Above blackish-brown, varying to black, the feathers usually bordered terminally with ochraceous or rusty; forehead usually more or less washed with the same. Beneath ochraceous, varying in shade; the whole surface with longitudinal stripes of blackish. Inner webs of tail-feathers and primaries with numerous transverse elliptical spots of ochraceous. Hab. Cosmopolitan.

a. Young dark brown above, the feathers bordered with rusty or whitish. Beneath white or ochraceous, with narrow longitudinal stripes of dusky. Inner webs of tail-feathers with transverse bars.

Auriculars white, cutting off the black of the cheeks with a prominent “mustache.”

Beneath pure white, the breast and middle of the abdomen without markings. Wing, 12.75; tail, 7.30; culmen, .80; tarsus, 2.00; middle toe, 1.80. Hab. Eastern Asia … var. orientalist.53

Beneath pale ochraceous, the breast always with longitudinal dashes, or elliptical spots, of dusky; middle of abdomen barred. Wing, 11.50–14.30; tail, 7.00–8.50; culmen, .72–.95; tarsus, 1.65–2.20; middle toe, 1.80–2.30. Hab. Europe … var. communis.54

Beneath varying from deep ochraceous to nearly pure white, the breast never with distinct longitudinal or other spots, usually with none at all. Middle of abdomen barred, or not. Wing, 11.30–14.75; tail, 6.00–9.00; culmen, .75–1.00; tarsus, 1.60–2.10; middle toe, 1.75–2.20. Hab. America (entire continent) … var. anatum.

Auriculars black, nearly, or quite, as far down as the lower end of the “mustache.”

Beneath varying from deep ochraceous to white, the breast streaked or not. Lower parts more uniformly and heavily barred than in the other races. Young with narrower streaks beneath. Wing, 11.15–12.60; tail, 6.11–8.00; culmen, .81–.90; tarsus, 1.60–2.05; middle toe, 1.75–2.15. Hab. Australia … var. melanogenys.55

b. Young unvariegated brownish-black above. Beneath brownish-black, faintly streaked with white, or nearly unvariegated. Inner webs of tail-feathers without transverse bars.

Wing, 14.90–15.09; tail, 8.50; culmen, .95–1.00; tarsus, 2.10; middle toe, 2.15–2.21. Hab. Northwest coast of North America, from Oregon to Sitka … var. pealei.

B. Second quill longest; first with inner web emarginated, the second with inner web not sinuated. Young without longitudinal stripes on lower parts. Adult and young stages hardly appreciably different.

Above plumbeous or black; beneath black from the jugulum to the tibiæ, with transverse bars of white, ochraceous, or rufous; throat and jugulum white, white and rufous, or wholly ochraceous, with a semicircular outline posteriorly; tibiæ, anal region, and crissum uniform deep rufous, or spotted with black on an ochraceous or a white and rufous ground. Adult. Plumbeous above, the feathers darker centrally, and with obscure darker bars posteriorly; jugulum immaculate. Young. Black above, the feathers bordered terminally with rusty, or else dark plumbeous without transverse bars; jugulum with longitudinal streaks.

2. F aurantius.56 Wing, 9.50–12.00; tail, 5.40–6.25; culmen, .96; tarsus, 1.50–1.60; middle toe, 1.75–2.10. Second quill longest; first longer than third. Crissum ochraceous, or white and rufous, with large transverse spots of black; upper tail-coverts sharply barred with pure white or pale ash. Adult. Above plumbeous-black, the feathers conspicuously bordered with plumbeous-blue. Throat and jugulum immaculate; white centrally and anteriorly, deep rufous laterally and posteriorly. Tibiæ plain rufous. Young. Above uniform dull black, the feathers sometimes bordered inconspicuously with rusty. Throat and jugulum varying from white to ochraceous or rufous (this always deepest laterally and posteriorly). Tibiæ sometimes thickly spotted transversely with black. Hab. Tropical America, north to Southern Mexico.

3. F. rufigularis.57 Wing, 7.20–9.00 (♂, wing, 7.70; tail, 3.95–5.50; culmen, .45–.58; tarsus, 1.20–1.55; middle toe, 1.15–1.40). Second quill longest; first longer than third. Crissum uniform deep reddish-rufous, rarely barred with white and dusky. Upper tail-coverts obsoletely barred with plumbeous.

Adult. Above plumbeous-black, the feathers lightening into plumbeous-blue on the edges and ends, and showing obscure bars on the posterior portions. Throat and jugulum ochraceous-white, the ochraceous tinge deepest posteriorly and without any streaks. Young. Above plumbeous-black, without lighter obscure bars, or with a brownish cast, and with faint rusty edges to the feathers. Throat and jugulum deep soft ochraceous, deepest laterally, the posterior portion usually with a few longitudinal streaks of dusky. Hab. Tropical America, north to Middle Mexico.

Falco communis, Gmel

Var. anatum, Bonap

AMERICAN PEREGRINE FALCON; DUCK HAWK

? Accipiter falco maculatus, Briss. Orn. I, 329. ? Falco nævius, Gmel. S. N. 1789, 271. Falco communis ζ, and F. communis η, Lath. Ind. Orn. p. 31. Falco communis, Coues, Key, 1872, 213, f. 141. Falco peregrinus, Ord. Wils. Am. Orn. 1808, pl. lxvi.—Sab. L. Trans. XII, 529.—Rich. Parry’s 2d Voy. App. 342.—Ib. F. B. A. II, 1831, 23.—Bonap. N. Y. Lyc. II, 27.—Ib. Isis, 1832, 1136; Consp. 1850, 23, No. 4.—King, Voy. Beag. I, 1839, 532.—James. Wils. Am. Orn. 677, Synop. 1852, 683.—Wedderb. Jard. Contr. to Orn. 1849, 81.—Woodh. Sitgr. Zuñi, 1853, 60.—Giraud, B. Long Island, 1844, 14.—Peale, U. S. Ex. Ex. 1848, 66.—Gray, List B. Brit. Mus. 1841, 51. Falco anatum, Bonap. Eur. & N. Am. B. 1838, 4.—Ib. Rev. Zoöl. 1850, 484.—Bridg. Proc. Zoöl. Soc. pl. xi, 109.—Ib. Ann. N. H. XIII, 499.—Gosse, B. Jam. 1847, 16.—Cass. B. Cal. & Tex. 1854, 86.—Ib. Birds N. Am. 1858, 7.—De Kay, Zoöl. N. Y. II, 13, pl. iii, f. 8.—Nutt. Man. 1833, 53.—Peab. B. Mass. 1841, 83.—Strickl. Orn. Syn. I, 1855, 83.—Blakist. Ibis, III, 1861, 315.—March, Pr. Ac. N. S. 1863, 304. Falco nigriceps, Cass. B. Cal. & Tex. I, 1853, 87.—Ib. Birds N. Am. 1858, 8.—Strickl. Orn. Syn. I, 85.—Coop. & Suckl. P. R. R. Rep’t, VII, ii, 1860, 142.—Gray, Hand List, I, 1869, 19, No. 166.—Sharpe, Ann. & Mag. N. H. Falco orientalis, (Gm.) Gray, Hand List, I, 1869, 19, No. 165 (in part). ? Falco cassini, Sharpe, Ann. & Mag. N. H.

Sp. Char. Adult (♂, 43,134, Fort Resolution, Brit. N. Am., June; J. Lockhart). Upper parts dark bluish-plumbeous, approaching black anteriorly, but on rump and upper tail-coverts becoming fine bluish plumbeous-ash. On the head and neck the continuous plumbeous-black covers all the former except the chin and throat, and the back portion of the latter; an invasion or indentation of the white of lower parts up behind the ear-coverts separating that of the cheeks from the posterior black, throwing the former into a prominent angular patch; forehead and lores grayish. All the feathers above (posterior to the nape) with transverse bars of plumbeous-black, these most sharply defined posteriorly, where the plumbeous is lightest. Tail black, more plumbeous basally, very faintly paler at the tip, and showing ten or eleven transverse narrow bands of plumbeous, these most distinct anteriorly; the bars are clearest on inner webs. Alula, primary and secondary coverts, secondaries and primaries, uniform plumbeous-black, narrowly whitish on terminal margin, most observable on secondaries and inner primaries. Lower parts white, tinged with delicate cream-color, this deepest on the abdomen; sides and tibiæ tinged with bluish. Chin, throat, and jugulum immaculate; the breast, however, with faint longitudinal shaft-streaks of black; sides, flanks, and tibiæ distinctly barred transversely with black, about four bars being on each feather; on the lower tail-coverts they are narrower and more distant; on the abdomen the markings are in the form of circular spots; anal region barred transversely. Lining of the wing (including all the under coverts) white tinged with blue, and barred like the sides; under surface of primaries slaty, with elliptical spots or bars of creamy-white on inner webs, twelve on the longest. Wing-formula, 2–1–3. Wing, 12.25; tail, 6.00; tarsus, 1.60; middle toe, 1.85; outer, 1.40; inner, 1.20; posterior, .80; culmen, .80.

♀ (13,077, Liberty Co., Georgia; Professor J. L. Leconte). Like the male, but ochraceous tinge beneath deeper; no ashy wash; bands on the tail more sharply defined, about ten dark ones being indicated; outer surface of primaries and secondaries with bands apparent; tail distinctly tipped with ochraceous-white. Inner web of longest primary with thirteen, more reddish, transverse spots. White of neck extending obliquely upward and forward toward the eye, giving the black cheek-patch more prominence. Markings beneath as in the male. Wing-formula the same. Wing, 14.50; tail, 7.00; tarsus, 1.95; middle toe, 2.10; culmen, .95.

Juv. (♂, 53,193, Truckee River, Nevada, July 24, 1867; R. Ridgway: first plumage). Above plumbeous-black, tail more slaty. Every feather broadly bordered terminally with dull cinnamon; these crescentic bars becoming gradually broader posteriorly, narrower and more obsolete on the head above. Tail distinctly tipped with pale cinnamon, the inner webs of feathers with obsolete transverse spots of the same, these touching neither the edge nor the shaft; scarcely apparent indications of corresponding spots on outer webs. Region round the eye, and broad “mustache” across the cheeks, pure black, the latter more conspicuous than in the older stages, being cut off posteriorly by the extension of the cream-color of the neck nearly to the eye. A broad stripe of pale ochraceous running from above the ear-coverts back to the occiput, where the two of opposite sides nearly meet. Lower parts purplish cream-color, or rosy ochraceous-white, deepest posteriorly; jugulum, breast, sides, flanks, and tibiæ with longitudinal stripes of plumbeous-black, these broadest on flanks and abdomen, and somewhat sagittate on the tibiæ; lower tail-coverts with distant transverse bars. Lining of the wing like the sides, but the markings more transverse; inner web of longest primary with nine transverse purplish-ochre spots. Wing-formula, 2–1, 3. Wing, 12.50; tail, 7.00. Length, 16.50; expanse, 39.25. Weight, 1½ lbs. Basal half of bill pale bluish-white, cere rather darker; terminal half (rather abruptly) slate-color, the tip deepening into black; iris very dark vivid vandyke-brown; naked orbital space pale bluish-white, with a slight greenish tint; tarsi and toes lemon-yellow, with a slight green cast; claws jet-black.

Hab. Entire continent of America, and neighboring islands.

Localities: Guatemala (Scl. Ibis I, 219); Veragua (Salv. P. Z. S. 1867, 158); Sta. Cruz (Newton, Ibis, I, 63); Trinidad (Taylor, Ibis, 1864, 80); Bahamas (Bryant, Pr. Bost. Soc. 1859, VII); Cuba (Cab. Journ. II, lxxxiii); (Gundl. Repert. 1865, 225); Jamaica, (Gosse, B. Jam. 16; March, Pr. Ac. N. S. 1863, 304, et Mus. S. I.); Tierra del Fuego (Sharpe, Ann. & Mag. N. H.; “F. cassini, Sharpe”).

The young plumage above described corresponds exactly with that of young peregrinus from Europe, a comparison of the specimen above described with one of the same age from Germany (54,064, Schlüter Col.) showing no differences that can be expressed. Many American specimens in this plumage (as 19,397, Fort Simpson) show a wash of whitish over the forehead and anterior part of the crown; having before us but the one specimen, we cannot say whether or not this is ever seen in the European bird. Specimens more advanced in season—perhaps in second year—are colored as follows: The black above is more brownish, the feathers margined with pale brown,—these margins broader, and approaching to white, on the upper tail-coverts; the tail shows the ochraceous bars only on inner webs. The supraoral stripe of the youngest plumage is also quite apparent.

A still younger one from the same locality (No. 37,397) has the upper plumage similar to the last, the pale edges to the feathers, however, more distinct; tail with conspicuous spots. White beneath clearer, and invading the dusky of the head above as far back as the middle of the crown; the supraoral stripe is distinct, scarcely interrupted across the nape.

In the adult plumage the principal variation is in the extent and disposition of the bars beneath. In most individuals they are regularly transverse only laterally and posteriorly, those on the belly being somewhat broken into more irregular cordate spots, though always transverse; in no American specimen, however, are they as continuously transverse as in a male (No. 18,804) from Europe, which, however, in this respect, we think, forms an exception to most European examples, at least to those in the Smithsonian Collection. All variations in the form, thickness, and continuity of the markings below, and in the distinctness of the bars above, are individual.

Very old males (as 49,790, Fort Yukon; 27,188, Moose Factory (type of Elliott’s figure of F. peregrinus, in Birds of America); and 42,997, Spanishtown, Jamaica) lack almost entirely the reddish tinge beneath, and have the lateral and posterior portions strongly tinged with blue; the latter feature is especially noticeable in the specimen from Jamaica, in which also the bars are almost utterly wanting medially. Immature birds from this island also lack to a great degree the ochraceous tinge, leaving the whitish everywhere purer.

A female adult European bird differs from the average of North American examples in the conspicuous longitudinal streaks on the jugulum; but in a male these are hardly more distinct than in 13,077, ♀, Liberty Co., Georgia; 11,983, “United States”; 35,456, Peel’s River; 35,449, ♀, and 35,445, ♀, Fort Yukon, Alaska; 35,452, La Pierre’s Hous., H. B. Ter.; 35,459 ♂, Fort Anderson; and 28,099 ♀, Hartford, Conn. In none of these, however, are they so numerous and conspicuous as in a European female from the Schlüter Collection, which, however, differs in these respects only from North American specimens.

A somewhat melanistic individual (in second year? 32,735, Chicago, Ill.; Robert Kennicott) differs as follows: Above continuously pure black; upper tail-coverts and longer scapulars bordered terminally with rusty-whitish. Tail distinctly tipped with white; the inner webs of feathers with eight elliptical transverse bars of pale ochraceous, and indications of corresponding spots of the same on outer webs, forming as many inconspicuous bands. Beneath ochraceous-white; the neck, breast, and abdomen thickly marked with broad longitudinal stripes of clear black,—those on the jugulum cuneate, and on the breast and abdomen broadly sagittate; the tibiæ with numerous cordate spots, and sides marked more transversely; lower tail-coverts with narrow distant transverse bars. On the chin and throat only, the whitish is immaculate, on the other portions being somewhat exceeded in amount by the black. Inner web of longest primary with seven transverse elliptical bars of cream-color. Wing, 12.20; tail, 9.40.

Whether the North American and European Peregrine Falcons are or are not distinct has been a question undecided up to the present day; almost every ornithologist having his own peculiar views upon the relationship of the different forms which have been from time to time characterized. The most favorably received opinion, however, seems to be that there are two species on the American continent, and that one of these, the northern one, is identical with the European bird. Both these views I hold to be entirely erroneous; for after examining and comparing critically a series of more than one hundred specimens of these birds, from every portion of America (except eastern South America), including nearly all the West India Islands, as well as numbers of localities throughout continental North and South America, I find that, with the exception of the melanistic littoral race of the northwest coast (var. pealei), they all fall under one race, which, though itself exceedingly variable, yet possesses characters whereby it may always be distinguished from the Peregrine of all portions of the Old World.

There is such a great amount of variability, in size, colors, and markings, that the F. nigriceps, Cassin, must be entirely ignored as being based upon specimens not distinguishable in any respect from typical anatum. Judging from the characters assigned to the F. cassini by its describer (who evidently had a very small series of American specimens at his command), the latter name must also most probably fall into the list of synonymes of anatum.

Slight as are the characters which separate the Peregrines of the New and Old World, i.e. the immaculate jugulum of the former and the streaked one of the latter, they are yet sufficiently constant to warrant their separation as geographical races of one species; along with which the F. melanogenys, Gould (Australia), F. minor, Bonap. (South Africa), F. orientalis, Gmel. (E. Asia), and F. calidus, Lath. (Southern India and East Indies), must also rank as simple geographical races of the same species. Whether the F. calidus is tenable, I am unable to state, for I have not seen it; but the others appear to be all sufficiently differentiated. The F. radama, Verreaux (Gray’s Hand List, p. 19, No. 170), Mr. Gurney writes me, is the young female of var. minor. Whether the F. peregrinator, Sundevall (Gray’s Hand List, No. 169), is another of the regional forms of F. communis, or a distinct species, I am not able at present to say, not having specimens accessible to me for examination.

Mr. Cassin’s type of “nigriceps” (13,856, ♂, July), from Chile, is before me, and upon comparison with adult males from Arctic America presents no tangible differences beyond its smaller size; the wing is a little more than half an inch, and the middle toe less than the eighth of an inch, shorter than in the smallest of the North American series,—a discrepancy slight indeed, and of little value as the sole specific character; the plumage being almost precisely similar to that of the specimen selected for the type of the description at the head of this article. In order to show the little consequence to be attached to the small size of the individual just mentioned, I would state that there is before me a young bird, received from the National Museum of Chile, and obtained in the vicinity of Santiago, which is precisely similar in plumage to the Nevada specimen described, and in size is even considerably larger, though it is but just to say that it is a female; the wing measures 13.25, instead of 12.50, and the middle toe, 2.00, instead of 1.85. No. 37,336, Tres Marias Islands, Western Mexico,—a young male in second year,—has the wing just the same length as in the smallest North American example, while in plumage it is precisely similar to 26,785, of the same age, from Jamaica. No. 4,367, from Puget’s Sound, Washington Territory,—also a young male,—has the wing of the same length as in the largest northern specimen, while the plumage is as usual.

Two adult females from Connecticut (Nos. 28,099 and 32,507, Talcott Mt.) are remarkable for their very deep colors, in which they differ from all other North American examples which I have seen, and answer in every particular to the description of F. cassini, Sharpe, above cited. The upper surface is plumbeous-black, becoming deep black anteriorly, the head without a single light feather in the black portions; the plumbeous bars are distinct only on the rump, upper tail-coverts, and tail, and are just perceptible on the secondaries. The lower parts are of a very deep reddish-ochraceous, deepest on the breast and abdomen, where it approaches a cinnamon tint,—the markings, however, as in other examples. They measure, wing, 14.75; tail, 7.50; culmen, 1.05–1.15; tarsus, 2.00; middle toe, 2.30. They were obtained from the nest, and kept in confinement three years, when they were sacrificed to science. The unusual size of the bill of these specimens (see measurements) is undoubtedly due to the influence of confinement, or the result of a modified mode of feeding. The specimens were presented by Dr. S. S. Moses, of Hartford.

An adult male (No. 8,501) from Shoal-water Bay, Washington Territory, is exactly of the size of the male described. In this specimen there is not the slightest creamy tinge beneath, while the blue tinge on the lower parts laterally and posteriorly is very strong. No. 52,818, an adult female from Mazatlan, Western Mexico, has the wing three quarters of an inch shorter than in the largest of four northern females, and of the same length as in the smallest; there is nothing unusual about its plumage, except that the bars beneath are sparse, and the ochraceous tinge quite deep. No. 27,057, Fort Good Hope, H. B. T., is, however, exactly similar, in these respects, and the wing is but half an inch longer. In No. 47,588, ♂, from the Farallones Islands, near San Francisco, California, the wing is the same length as in the average of northern and eastern specimens, while the streaks on the jugulum are nearly as conspicuous as in a male from Europe.

In conclusion, I would say that the sole distinguishing character between the Peregrines from America and those from Europe, that can be relied on, appears to be found in the markings on the breast in the adult plumage; in all the specimens and figures of var. communis that I have seen, the breast has the longitudinal dashes very conspicuous; while, as a general rule, in anatum these markings are entirely absent, though sometimes present, and occasionally nearly as distinct as in European examples. Therefore, if this conspicuous streaking of the breast is found in all European specimens, the American bird is entitled to separation as a variety; but if the breast is ever immaculate in European examples, then anatum must sink into a pure synonyme of communis. The var. melanogenys is distinguished from both communis and anatum by the black auriculars, or by a greater amount of black on the side of the neck, and by more numerous and narrower bars on the under surface. In the former feature examples of anatum from the southern extremity of South America approach quite closely to the Australian form, as might be expected from the relative geographical position of the two regions. The var. minor is merely the smaller intertropical race of the Old World, perhaps better characterized than the tropical American form named F. nigriceps by Cassin, the characters of which are so unimportant, and withal so inconstant, as to forbid our recognizing it as a race of the same rank with the others.

LIST OF SPECIMENS EXAMINED

National Museum, 45; Boston Society, 4; Philadelphia Academy, 22; Museum Comp. Zoöl. 5; New York Museum, 3; G. N. Lawrence, 6; R. Ridgway, 3. Total, 88.

Var. pealei, Ridgway

BLACK PEREGRINE FALCON

? ? Accipiter falco niger, Briss. Orn. I, 337. ? ? Falco niger, Gmel. S. N. 1789, 270. Falco polyagrus, Cass. B. Cal. & Tex. pl. xvi (dark figure).

Sp. Char. In colors almost exactly similar to F. gyrfalco, var. labradora. Above continuous dark vandyke-brown, approaching brownish-black on the head, which is variegated only on the gular region, and inclining to grayish-brown on the tail; the whole surface entirely free from spots or markings of any kind. Beneath similar in color to the upper parts, but the feathers edged with whitish, this rather predominating on the throat; flanks and tibiæ with roundish white spots; lower tail-coverts with broad transverse bars of white. Lining of the wing with feathers narrowly tipped with white; inner webs of primaries with narrow, transverse elliptical spots of cream-color; inner webs of tail-feathers with badly defined, irregular, similar spots, or else with these wanting, the whole web being plain dusky-brown.

No. 12,022 (♀, Oregon; T. R. Peale). Wing, 15.00; tail, 8.50; culmen, .95; tarsus, 2.10; middle toe, 2.15. (Figured by Cassin as F. polyagrus, in Birds of California and Texas, pl. xvi.)

No. 45,814 (♀, Sitka, Alaska, May, 1866; F. Bischoff). Wing, 14.90; tail, 8.50; tarsus, 2.10; middle toe, 2.20. The two similar in color, but in the latter the white streaks on the lower parts a little broader, and the middle of the auriculars slightly streaked.

Hab. Northwest coast of North America, from Oregon to Sitka.

This curious race of Falco communis is a good illustration of the climatic peculiarity of the northwest coast region, to which I have often referred before; the same melanistic tendency being apparent in birds of other species from the same region, as an example of which I may mention the Black Merlin (Falco æsalon, var. suckleyi), which is a perfect miniature of the present bird.

Habits. The Great-footed Hawk of North America is very closely allied to the well-known Peregrine Falcon of Europe, and so closely resembles it that by many writers, even at the present day, it is regarded as identical with it. Without doubt, the habits of the two races are very nearly the same, though the peculiarities of the North American bird are not so well known as are those of the European. In its distribution it is somewhat erratic, for the most part confined to the rocky sea-coast, the river-banks, and the high ground of the northeastern parts of America. It is known to breed in a few isolated rocky crags in various parts of the country, even as far to the south as Pennsylvania, and it occurs probably both as migrant and resident in several of the West India Islands, in Central and in South America. A single specimen was taken by Dr. Woodhouse in the Creek country of the Indian Territory. Two individuals are reported by Von Pelzeln as having been taken in Brazil. The Newtons met with it in St. Croix. Mr. Gosse found it in Jamaica, and Dr. Gundlach gives it as a bird of Cuba. Jardine states it to be a bird of Bermuda, and also that it has been taken in the Straits of Magellan. A single specimen was taken at Dueñas, Guatemala, in February, by Mr. Salvin.

On the Pacific coast this Falcon has been traced as far south as the limit of the land. Dr. Cooper met with only two pairs, in March, 1854, frequenting a high wooded cliff at Shoal-water Bay. Dr. Suckley procured a single specimen from Steilacoom. Dr. Cooper states that the habits of these corresponded with those described for the F. anatum and F. peregrinus, and that, like these Falcons, it is a terror to all land animals weaker than itself. It is said to breed on the rocky cliffs of the Pacific.

An individual of this bird was taken by Colonel Grayson at the Tres Marias Islands. When shot, it was endeavoring to capture a Sparrow-hawk, indicating its indifference as to the game it pursues. He adds that this bird attacks with vigor everything it sees, from the size of a Mallard Duck down, and is the terror of all small birds. Its range must be very great, as it often ventures far out to sea. On his passage from Mazatlan to San Francisco, in 1858, on the bark Carlota, one of these Falcons came on board more than a hundred miles off the coast of Lower California, and took up its quarters on the main-top yard, where it remained two days, during which time it captured several Dusky Petrels. It would dart headlong upon these unsuspecting birds, seldom missing its aim. It would then return to its resting-place and partly devour its prize. At other times it dropped its victims into the sea in wanton sport. Finally, as if tired of this kind of game, it made several wide circles around the ship, ascended to a considerable height, and departed in the direction of the Mexican shore.

This Falcon is found along the Atlantic coast from Maine to the extreme northern portion, breeding on the high rocky cliffs of Grand Menan and in various favorable situations thence northward. A few breed on Mount Tom, near the Connecticut River in Massachusetts, on Talcott Mountain in Connecticut, in Pennsylvania, and near Harper’s Ferry, in Maryland.

Mr. Boardman has several times taken their eggs from the cliffs of Grand Menan, where they breed in April, or early in May. In one instance he found the nest in close proximity to that of a pair of Ravens, the two families being apparently on terms of amity or mutual tolerance.

For several years two or more pairs of these birds have been known to breed regularly on Mount Tom, near Northampton. The nests were placed on the edges of precipitous rocks very early in the spring, the young having been fully grown by the last of June. Their young and their eggs have been taken year after year, yet at the last accounts they still continued to nest in that locality. Dr. W. Wood has also found this species breeding on Talcott Mountain, near Hartford. Four young were found, nearly fledged, June 1. In one instance four eggs were taken from a nest on Mount Tom, by Mr. C. W. Bennett, as early as April 19. This was in 1864. Several times since he has taken their eggs from the same eyrie, though the Hawks have at times deserted it and sought other retreats. In one year a pair was twice robbed, and, as is supposed, made a third nest, and had unfledged young as late as August. Mr. Allen states that these Hawks repair to Mount Tom very early in the spring, and carefully watch and defend their eyrie, manifesting even more alarm at this early period, when it is approached, than they evince later, when it contains eggs or young. Mr. Bennett speaks of the nest as a mere apology for one.

This Hawk formerly nested on a high cliff near the house of Professor S. S. Haldeman, Columbia, Penn., who several times procured young birds which had fallen from the nest. The birds remained about this cliff ten or eleven months of the year, only disappearing during the coldest weather, and returning with the first favorable change. They bred early in spring, the young leaving the nest perhaps in May. Professor Haldeman was of the opinion that but a single pair remained, the young disappearing in the course of the season.

Sir John Richardson, in his Arctic expedition in 1845, while descending the Mackenzie River, latitude 65°, noticed what he presumed to be a nest of this species, placed on the cliff of a sandstone rock. This Falcon was rare on that river.

Mr. MacFarlane found this species not uncommon on the banks of Lockhart and Anderson Rivers, in the Arctic regions. In one instance he mentions finding a nest on a cliff thirty feet from the ground. There were four eggs lying on a ledge of the shale of which the cliff was composed. Both parents were present, and kept up a continued screaming, though at too great a distance for him to shoot either. He adds that this bird is by no means scarce on Lockhart River, and he was informed that it also nests along the ramparts and other steep banks of the Upper Anderson, though he has not been able to learn that it has been found north of Fort Anderson. In another instance the nest was on a ledge of clayey mud,—the eggs, in fact, lying on the bare ground, and nothing resembling a nest to be seen. A third nest was found on a ledge of crumbling shale, along the banks of the Anderson River, near the outlet of the Lockhart. This Hawk, he remarks, so far as he was able to observe, constructs no nest whatever. At least, on the Anderson River, where he found it tolerably abundant, it was found to invariably lay its eggs on a ledge of rock or shale, without making use of any accessory lining or protection, always availing itself of the most inaccessible ledges. He was of the opinion that they do not breed to the northward of the 68th parallel. They were also to be found nesting in occasional pairs along the lime and sandstone banks of the Mackenzie, where early in August, for several successive years, he noticed the young of the season fully fledged, though still attended by the parent birds.

In subsequent notes, Mr. MacFarlane repeats his observations that this species constructs no nest, merely laying its eggs on a ledge of shale or other rock. Both parents were invariably seen about the spot. In some instances the eggs found were much larger than in others.

Mr. Dall mentions shooting a pair near Nuk´koh, on the Yukon River, that had a nest on a dead spruce. The young, on the 1st of June, were nearly ready to fly. It was not a common species, but was found from Nulato to Sitka and Kodiak.

In regard to general characteristics of this Falcon, they do not apparently differ in any essential respects from those of the better-known Falco communis of the Old World. It flies with immense rapidity, rarely sails in the manner of other Hawks, and then only for brief periods and when disappointed in some attempt upon its prey. In such cases, Mr. Audubon states, it merely rises in a broad spiral circuit, in order to reconnoitre a space below. It then flies swiftly off in quest of plunder. These flights are made in the manner of the Wild Pigeon. When it perceives its object, it increases the flappings of its wings, and pursues its victim with a surprising rapidity. It turns, and winds, and follows every change of motion of the object of pursuit with instantaneous quickness. Occasionally it seizes a bird too heavy to be managed, and if this be over the water it drops it, if the distance to land be too great, and flies off in pursuit of another. Mr. Audubon has known one of this species to come at the report of a gun, and carry off a Teal not thirty steps distant from the sportsman who had killed it. This daring conduct is a characteristic trait.

This bird is noted for its predatory attacks upon water-fowl, but it does not confine itself to such prey. In the interior, Richardson states that it preys upon the Wild Pigeon, and upon smaller birds. In one instance Audubon has known one to follow a tame Pigeon to its house, entering it at one hole and instantly flying out at the other. The same writer states that he has seen this bird feeding on dead fish that had floated to the banks of the Mississippi. Occasionally it alights on the dead branch of a tree in the neighborhood of marshy ground, and watches, apparently surveying, piece by piece, every portion of the territory. As soon as it perceives a suitable victim, it darts upon it like an arrow. While feeding, it is said to be very cleanly, tearing the flesh, after removing the feathers, into small pieces, and swallowing them one by one.

The European species, as is well known, was once largely trained for the chase, and even to this day is occasionally used for this purpose; its docility in confinement, and its wonderful powers of flight, rendering it an efficient assistant to the huntsman. We have no reason to doubt that our own bird might be made equally serviceable.

Excepting during the breeding-season, it is a solitary bird. It mates early in February, and even earlier in the winter. Early in the fall the families separate, and each bird seems to keep to itself until the period of reproduction returns.

In confinement, birds of this family become quite tame, can be trained to habits of wonderful docility and obedience, and evince even an affection for the one who cares for their wants.

This species appears to nest almost exclusively on cliffs, and rarely, if ever, to make any nests in other situations. In a few rare and exceptional cases this Falcon has been known to construct a nest in trees. Mr. Ord speaks of its thus nesting among the cedar swamps of New Jersey; but this fact has been discredited, and there has been no recent evidence of its thus breeding in that State. Mr. Dall found its nest in a tree in Alaska, but makes no mention of its peculiarities.

The eggs of this species are of a rounded-oval shape, and range from 2.00 to 2.22 inches in length, and from 1.60 to 1.90 in width. Five eggs, from Anderson River, have an average size of 2.09 by 1.65 inches. An egg from Mount Tom, Mass., is larger than any other I have seen, measuring 2.22 inches in length by 1.70 in breadth, and differs in the brighter coloring and a larger proportion of red in its markings. The ground is a deep cream-color, but is rarely visible, being generally so entirely overlaid by markings as nowhere to appear. In many the ground-color appears to have a reddish tinge, probably due to the brown markings which so nearly conceal it. In others, nothing appears but a deep coating of dark ferruginous or chocolate-brown, not homogeneous, but of varying depth of coloring, and here and there deepening into almost blackness. In one egg, from Anderson River, the cream-colored ground is very apparent, and only sparingly marked with blotches of a light brown, with a shading of bronze. An egg from the cabinet of Mr. Dickinson, of Springfield, taken on Mount Tom, Massachusetts, is boldly blotched with markings of a bright chestnut-brown, varying greatly in its shadings.

Subgenus ÆSALON, Kaup

Æsalon, Kaup, 1829. (Type, Falco æsalon, Gmelin, = F. lithofalco, Gm.)

Hypotriorchis, Auct. nec Boie, 1826, the type of which is Falco subbuteo, Linn.

Dendrofalco, Gray, 1840. (Type, F. æsalon, Gmel.)

This subgenus contains, apparently, but the single species F. lithofalco, which is found nearly throughout the Northern Hemisphere, and in different climatic regions is modified into geographical races. Of these, North America possesses three, and Europe one; they may be distinguished as follows:—

Species and Races

F. lithofalco. Second and third quills longest; first usually shorter than, occasionally equal to, or rarely longer than, the fourth. Adult female, and young of both sexes. Above brownish, varying from pale earth-brown, or umber, to nearly black, plain, or with obscure transverse spotting of lighter; tail with five to eight lighter bands, which, however, are sometimes obsolete, except the terminal one. Beneath ochraceous-white, longitudinally striped with brown or dusky over the whole surface. Adult male (except in var. suckleyi and richardsoni?). Above plumbeous-blue, with darker shaft-streaks; tail with more or less distinct bands of black, and paler tip. Beneath much as in the female and young, but stripes usually narrower and more reddish. Wing, 7.20–9.00; tail, 4.90–6.30; culmen, .45–.60; tarsus, 1.30–1.60; middle toe, 1.15–1.51.

a. Adult male plumbeous-blue above; sexes very unlike in adult dress. Female and young without transverse spotting on upper parts.

Adult male. Tail deep plumbeous, tipped with ash, with six transverse series of dusky spots (which do not touch the shaft nor edge of the feathers) anterior to the subterminal zone, the black of which extends forward along the edge of the feather. Inner web of the longest primary with ten transverse spots of white. Streaks on the cheeks enlarged and blended, forming a conspicuous “mustache.” Pectoral markings linear black. The ochraceous wash deepest across the nape and breast, and along the sides, and very pale on the tibiæ. Adult female. Above brownish-plumbeous, the feathers becoming paler toward their margins, and with conspicuous black shaft-streaks. Tail with eight (three concealed) narrow bands of pale fulvous-ashy; longest primary with ten light spots on inner web. Outer webs of primaries with a few spots of ochraceous. Young. Similar to the ♀ adult, but with a more rusty cast to the plumage, and with more or less distinct transverse spots of paler on the upper parts. Wing, 7.60–9.00; tail, 5.10–6.30; culmen, .45–.55; tarsus, 1.35–1.47; middle toe, 1.15–1.35. Hab. Europe … var. lithofalco.58

Adult male. Tail light ash, tipped with white, and crossed by three or four nearly continuous narrow bands of black (extending over both webs, and crossing the shaft), anterior to the broad subterminal zone, the black of which does not run forward along the edge of the feathers. Inner web of longest primary with seven to nine transverse spots of white. Streaks on the cheeks sparse and fine, not condensed into a “mustache.” Pectoral markings broad clear brown. Ochraceous wash weak across the nape and breast, and along sides, and very deep on the tibiæ. Adult female. Above plumbeous-umber, without rusty margins to the feathers, and without conspicuous black shaft-streaks. Tail with only five (one concealed) narrow bands of pale ochraceous; outer webs of primaries without ochraceous spots; inner web of outer primary with eight spots of white. Young. Like the adult female, but darker. Wing, 7.90–8.25; tail, 5.15–5.25; tarsus, 1.00; middle toe, 1.25. Hab. Entire continent of North America; West Indies … var. columbarius.

b. Adult male not bluish? sexes similar? upper parts with lighter transverse spots.

Adult. Above light grayish-umber, or earth-brown, with more or less distinct lighter transverse spots; secondaries crossed by three bands of ochraceous spots, and outer webs of inner primaries usually with spots of the same. Tail invariably with six complete and continuous narrow bands of dull white. Beneath white, with broad longitudinal markings of light brown, these finer and hair-like on the tibiæ and cheeks, where they are sparse and scattered, not forming a “mustache.” Top of the head much lighter than the back. Young. Similar, but much tinged with rusty above, all the white portions inclining to pale ochraceous. Wing, 7.70–9.00; tail, 5.00–6.30; culmen, .50–.60; tarsus, 1.40–1.65; middle toe, 1.20–1.51. Second and third quills longest; first equal to fourth, slightly shorter, or sometimes slightly longer. Hab. Interior plains of North America, between the Mississippi River and the Rocky Mountains, from the Arctic regions to Texas … var. (?) richardsoni.

c. Adult male not bluish? sexes similar? upper parts without transverse spots, and tail without lighter bands, except at the tip.

Above plain brownish-black; the tail narrowly tipped with whitish, but without other markings; inner webs of the primaries without lighter spots. Beneath pale ochraceous broadly striped with sooty-black. Wing, 7.35–8.50; tail, 5.25–5.75; culmen, .50–.55; tarsus, 1.30–1.62; middle toe, 1.25–1.35. Hab. Northwest coast region from Oregon to Sitka … var. suckleyi.

Falco (Æsalon) lithofalco (Gmelin)

Var. columbarius, Linnæus

PIGEON HAWK; AMERICAN MERLIN

Falco columbarius, Linn. Syst. Nat. 1766, p. 128.—Gmel. Syst. Nat. 1789, p. 281.—Lath. Ind. Orn. I, 44, 1790; Syn. I, 101, sp. 86; Supp. I, 27, 1802; Gen. Hist. I, 278, 1821.—Daud. Tr. Orn. II, 83, 1800.—Shaw. Zoöl. VII, 188, 1812.—Wils. Am. Orn. pl. xv, fig. 3, 1808.—Jard. (Wils.) Am. Orn. I, p. 254, 1808.—James. (Wils.) Am. Orn. I, 61.—Brew. (Wils.) Am. Orn. I, 683, 1852.—Rich. Faun. Bor. Am. II, 35, 1831.—Aud. Syn. B. A. p. 16, 1839; Orn. Biog. I, 466.—Bonap. Ann. Lyc. N. Y. II, 28; Isis, 1832, p. 1136; Eur. & N. Am. B. p. 4, 1838.—Nutt. Man. I, 60, 1833.—Cuv. Règ. An. (ed. 2), I, 322, 1829.—Less. Tr. Orn. p. 92, 1831.—Forst. Phil. Trans. LXII, 382, 1772.—Swains. Classif. B. II, p. 212, 1837.—Jard. Ann. Nat. Hist. XVIII, 118.—Gosse, B. Jam. p. 17, 1847.—Sagra, Hist. Nat. Cuba Ois. p. 23.—Wedderb. Jard. Cont. Orn. 1849, p. 81.—Hurdis, Jard. Cont. Orn. 1850, p. 6.—De Kay, Zoöl. N. Y. II, 15, pl. iv, f. 9, 1844.—Giraud, B. Long Isl. p. 17.—Blackist. Ibis, III, 315. Tinnunculus columbarius, Vieill. Ois. Am. Sept. I, pl. xi, 1807; Nouv. Dict. Hist. Nat. XII, 104, 1819; Enc. Méth. III, 1236, 1823. Hypotriorchis columbarius, Gray, List B. B. Mus. p. 55, 1844; Gen. B. fol. sp. 11, 1844.—Cass. B. Calif. & Tex. p. 90, 1854.—Woodh. (Sitg.) Exp. Zuñi & Colorad. p. 60, 1853.—Heerm. P. R. R. Rept. II, 31, 1855.—Newb. P. R. Rept. VI, 74, 1857.—Cass. B. N. Am. p. 9, 1858.—Cooper & Suck. P. R. R. Rept. XII, 1860, 142.—Coues, Pr. A. N. S. Phil. 1866, 6.—Brewer, Oölogy, 12. Lithofalco columbarius, Bonap. Consp. Av. p. 26, 1850. Æsalon columbarius, Kaup, Monog. Falc. Cont. Orn. p. 54, 1850.—Gray, Hand List, I, 21, 1869. Falco obscurus, Gmel. Syst. Nat. p. 281, 1789.—Lath. Ind. Orn. p. 44, 1790; Syn. Supp. I, 38, 1802; Gen. Hist. I, 272, 1821.—Daud. Tr. Orn. II, p. 123, 1800. Falco intermixtus, Daud. Tr. Orn. II, p. 141, 1800.—Lath. Gen. Hist. I, 136, 1821. F. temerarius, Aud. B. Am. pls. lxxv, xcii, 1831; Orn. Biog. I, 380, 1831. F. auduboni, Blackw. Res. Zoöl. 1840. Accipiter palumbarius, Catesb. Carol. I, pl. iii, 1754.

Sp. Char. Adult male. Above cinereous, varying in shade, but generally of a slaty-bluish cast; each feather with a distinct shaft-streak of black, these lines most conspicuous on the head above. Tail with a very broad subterminal band of black, about one inch in width; there are indications of three other bands, their continuity and distinction varying with the individual, but generally quite conspicuous, and each about half the width of the terminal one; the subterminal black band is succeeded by a terminal one of white, of about three-sixteenths of an inch in width, sometimes broader; on the lateral feathers the black bands are always conspicuous, being in form of transverse oblong spots, crossing the shaft, but less extended on the outer web, which is often immaculate except at the end, the broad terminal band always extending to the edge of the feather. Primaries dusky-black, margined terminally more or less distinctly with whitish (sometimes fading on the edge only); on the inner web is a series of about eight transverse oval spots of white, and generally corresponding to these are indications of bluish-ashy spots on the outer web. Beneath white, this purest on the throat, which is immaculate: there is generally a more or less strong tinge of fulvous beneath, this always prevalent on the tibiæ, and on a distinct collar extending round the nape, interrupting the blue above; the tibiæ frequently incline to ochraceous-rufous. Lateral portions of the head with fine streaks of dusky, these thickest on upper edge of the ear-coverts, leaving a distinct whitish superciliary streak, those of opposite sides meeting on the forehead. Breast, upper part of the abdomen, sides, and flanks, with longitudinal stripes of umber, each with a shaft-streak of black; on the flanks their shape is modified, here taking the form of spots running in chain-like series; tibiæ with narrower and darker streaks; lower tail-coverts with narrow central streaks like those on the tibiæ. Frequently there is a strong bluish shade on flanks and lower tail-coverts, sometimes replacing the brown of the spots on the former, and clouding in a similar form the latter. Length, 11.00; extent, 23.75; wing, 7.75.

Adult female. Pattern of coloration as in the male, but the colors different. The blue above replaced by dark umber-brown with a plumbeous cast, and showing more or less distinct darker shaft-lines; these on the head above very broad, giving a streaked appearance; white spots on inner webs of primaries more ochraceous than in the male. Tail dark plumbeous-brown, shading into blackish toward end, with five rather narrow ochraceous or soiled white bars, the first of which is concealed by the upper coverts, the last terminal. White beneath, less tinged with reddish than in the male, the tibiæ not different from the other portions; markings beneath as in the male.

Juv. Above plumbeous-brown, tinged with fulvous on head, and more or less washed with the same on the rump; frequently the feathers of the back, rump, scapulars, and wings pass into a reddish tinge at the edge; this color is, however, always prevalent on the head, which is conspicuously streaked with dusky. Tail plumbeous-dusky, darker terminally, with five regular light bars, those toward the base ashy, as they approach the end becoming more ochraceous; these bars are more continuous and regular than in the adult female, and are even conspicuous on the middle feathers. Primaries dusky, passing on edge (terminally) into lighter; spots on the inner webs broader than in the female, and pinkish-ochre; outer webs with less conspicuous corresponding spots of the same. Beneath soft ochraceous; spots as in adult female, but less sharply defined; tibiæ not darker than abdomen.

Hab. Entire continent of North America, south to Venezuela and Ecuador; West India Islands.

Localities: Ecuador (high regions in winter, Scl. P. Z. S. 1858, 451); Cuba (Cab. Jour. II, lxxxiii, Gundlach, Sept. 1865, 225); Tobago (Jard. Ann. Mag. 116); S. Texas (Dresser, Ibis, 1865, 323, breeding?); W. Arizona (Coues, Pr. A. N. S. 1866, 42); Costa Rica (Lawr. IX, 134); Venezuela (Scl. & Salv. 1869, 252).

LIST OF SPECIMENS EXAMINED

National Museum, 42; Boston Society, 11; Philadelphia Academy, 10; Museum Comp. Zoöl., 7; New York Museum, 3; G. N. Lawrence, 2; R. Ridgway, 4. Total, 79.

The plumage of the adult male, which is not as often seen as that of the younger stages and adult female, is represented in the Smithsonian Collection by fifteen specimens, from various parts of North America. Of these, an example from Jamaica exhibits the purest shades of color, though agreeing closely with some specimens from the interior of the United States; the cinereous above being very fine, and of a light bluish cast. The upper tail-coverts are tipped with white; the tail is a quarter of an inch longer than in any North American specimen, one half-inch longer than the average; the wing, however, is about the same.

A specimen from Santa Clara, California (4,475, Dr. J. G. Cooper), like most of those from the Pacific coast, has the cinereous very dark above, while beneath the ochraceous is everywhere prevalent; the flanks are strongly tinged with blue; the black bars of the tail are much broken and irregular. A specimen from Jamaica (24,309, Spanish Town; W. T. March), however, is even darker than this one, the stripes beneath being almost pure black; on the tail black prevails, although the bands are very regular. Nos. 27,061, Fort Good Hope, British America, 43,136, Fort Yukon, Alaska, and 51,305, Mazatlan, Mexico, have the streaks beneath narrow and linear; the ochraceous confined to the tibiæ, which are of a deep shade of this color.

Falco columbarius.

A specimen from Nicaragua (No. 40,957, Chinandega) is like North American examples, but the reddish tinge beneath is scarcely discernible, and confined to the tibiæ, which are but faintly ochraceous; the markings beneath are broad and deep umber, the black shaft-streak distinct.

In the adult female there is as little variation as in the male in plumage, the shade of brown above varying slightly, also the yellowish tinge beneath; the bars on the tail differ in continuity and tint in various specimens, although they are always five in number,—the first concealed by the coverts, the last terminal. In 19,382, Fort Simpson, British America, and 2,706, Yukon, R. Am. (probably very old birds), the light bars are continuous and pale dull ashy.

The young vary about the same as adults. Nos. 19,381, Big Island, Great Slave Lake; 5,483, Petaluma, California; and 3,760, Racine, Wisconsin,—are young males moulting, scattered feathers appearing on the upper parts indicating the future blue plumage.

Var. suckleyi, Ridgway

BLACK MERLIN

Sp. Char. A miniature of F. peregrinus, var. pealei. Above, uniform fuliginous-black, the secondaries and tail-feathers very narrowly but sharply tipped with white, and the primaries passing into whitish on their terminal margin; nuchal region with concealed spotting of pale rusty or dingy whitish. Beneath, longitudinally striped with fuliginous-black, or dark sooty-brown, and pale ochraceous; the former predominating on the breast, the latter prevailing on the throat and anal region. Sides and flanks nearly uniform dusky, with roundish white spots on both webs; lower tail-coverts with a broad sagittate spot of dusky on each feather. Lining of the wing fuliginous-dusky, with sparse, small roundish spots of white. Inner webs of primaries plain dusky, without spots, or else with them only faintly indicated. Tail plain dusky-black, narrowly tipped with white, and without any bands, or else with them only faintly indicated.

Male (No. 4,477, Shoalwater Bay, Washington Territory; J. G. Cooper). Wing, 7.35; tail, 5.25; culmen, .50; tarsus, 1.30; middle toe, 1.25.

Female (No. 5,832, Fort Steilacoom, Washington Territory, September, 1856; Dr. George Suckley). Wing, 8.50; tail, 5.70; culmen, .55; tarsus, 1.62; middle toe, 1.35.

Hab. Coast region of Northern California, Oregon, and Washington Territory (probably northward to Alaska). Puget Sound, Steilacoom, Yreka, California (Oct.), and Shoalwater Bay (National Museum).

The plumage of this race is the chief point wherein it differs from the other forms of the species; and in its peculiarities we find just what should be expected from the Oregon region, merely representing as it does the melanistic condition so frequently observable in birds from the northwest coast.

The upper parts are unicolored, being continuous blackish-plumbeous from head to tail. The tail is tipped with white, but the bars are very faintly indicated, being in No. 4,499 altogether wanting, while in 21,333 they can scarcely be discovered, and only four are indicated; in the others there is the usual number, but they are very obsolete. In No. 4,499, the most extreme example, the spots on the inner webs of the primaries are also wanting; the sides of the head are very thickly streaked, the black predominating, leaving the superciliary stripe ill-defined; the throat is streaked, and the other dark markings beneath are so exaggerated that they cover all portions, and give the prevailing color; the under tail-coverts have broad central cordate black spots.

Another specimen from this region (4,476, Puget Sound) is similar, but the spots on primaries are conspicuous, as in examples of the typical style; indeed, except in the most extreme cases, these spots will always be found indicated, leading us to the unavoidable conclusion that the specimens in question represent merely the fuliginous condition of the common species; not the condition of melanism, but the peculiar darkened plumage characteristic of many birds of the northwest coast, the habitat of the present bird; it should then be considered as rather a geographical race, co-equal to the Falco gyrfalco, var. labradora, F. peregrinus, var. pealei, and other forms, and not confounded with the individual condition of melanism, as seen in certain species of Buteones.

LIST OF SPECIMENS EXAMINED

National Museum, 6.

Second quill longest; first quill equal to, a little shorter than, or a little longer than, the fourth.

Var. richardsoni, Ridgway

RICHARDSON’S MERLIN

Falco æsalon, Rich. & Swains. F. B. A. II, pl. xxv, 1831.—Nutt. Man. Orn. II, 558.—Coues, P. A. N. S. Philad. 1866, p. 42 (in text). Falco (Hypotriorchis) richardsoni, Ridgway, P. A. N. S. Philad. Dec. 1870, 145. Falco richardsoni, Coues, Key, 1872, p. 214.

Sp. Char. Adult male like the female and young? The known stages of plumage more like the adult female and young of var. lithofalco (F. æsalon, Auct.) than like var. columbarius.

Adult male (Smithsonian, No. 5,171, mouth of the Vermilion River, near the Missouri, October 25, 1856; Lieutenant Warren, Dr. Hayden). Upper plumage dull earth-brown, each feather grayish-umber centrally, and with a conspicuous black shaft-line. Head above approaching ashy-white anteriorly, the black shaft-streaks being very conspicuous. Secondaries, primary coverts, and primaries margined terminally with dull white; the primary coverts with two transverse series of pale ochraceous spots; outer webs of primaries with spots of the same, corresponding with those on the inner webs. Upper tail-coverts tipped, and spotted beneath the surface, with white. Tail clear drab, much lighter than the primaries, but growing darker terminally, having basally a slightly ashy cast; crossed with six sharply defined, perfectly continuous bands (the last terminal) of ashy-white. Head, frontally, laterally, and beneath,—a collar around the nape (interrupting the brown above),—and the entire lower parts, white, somewhat ochraceous, this most perceptible on the tibiæ; cheeks and ear-coverts with sparse, fine hair-like streaks of black; nuchal collar, jugulum, breast, abdomen, sides, and flanks with a medial linear stripe of clear ochre-brown on each feather; these stripes broadest on the flanks; each stripe with a conspicuously black shaft-streak; tibiæ and lower tail-coverts with fine shaft-streaks of brown, like the broader stripes of the other portions. Chin and throat, only, immaculate. Lining of the wing spotted with ochraceous-white and brown, in about equal amount, the former in spots approaching the shaft. Inner webs of primaries with transverse broad bars of pale ochraceous,—eight on the longest. Wing-formula, 2, 3–4, 1. Wing, 7.70; tail, 5.00; culmen, .50; tarsus, 1.30; middle toe, 1.25; outer, .85; inner, .70; posterior, .50.

Adult female (58,983, Berthoud’s Pass, Rocky Mountains, Colorado Territory; Dr. F. V. Hayden, James Stevenson). Differing in coloration from the male only in the points of detail. Ground-color of the upper parts clear grayish-drab, the feathers with conspicuously black shafts; all the feathers with pairs of rather indistinct rounded ochraceous spots, these most conspicuous on the wings and scapulars. Secondaries crossed with three bands of deeper, more reddish ochraceous. Bands of the tail pure white. In other respects exactly as in the male. Wing-formula, 3, 2–4–1. Wing, 9.00; tail, 6.10; culmen, .55; tarsus, 1.40; middle toe, 1.51.

Young male (40,516, Fort Rice, Dacotah, July 20, 1865; Brig.-Gen. Alfred Sully, U. S. A., S. M. Rothammer). Differing from the adult only in minute details. Upper surface with the rusty borders of the feathers more washed over the general surface; the rusty-ochraceous forms the ground-color of the head,—paler anteriorly, where the black shaft-streaks are very conspicuous; spots on the primary coverts and primaries deep reddish-ochraceous; tail-bands broader than in the adult, and more reddish; the terminal one twice as broad as the rest (.40 of an inch), and almost cream-color in tint. Beneath pale ochraceous, this deepest on the breast and sides; markings as in the adult, but anal region and lower tail-coverts immaculate; the shaft-streaks on the tibiæ, also, scarcely discernible. Wing, 7.00; tail, 4.60.

Hab. Interior regions of North America, between the Mississippi Valley and the Rocky Mountains, from Texas to the Arctic regions.

LIST OF SPECIMENS EXAMINED

National Museum, 10; Museum Comp. Zoöl., 2; R. Ridgway, 3. Total, 15.

Since originally describing this bird, I have seen additional examples, and still consider it as an easily recognized race, not at all difficult to distinguish from columbarius. Now, however, I incline strongly to the theory that it represents merely the light form of the central prairie regions, of the common species; since its characters seem to be so analogous to those of the races of Buteo borealis and Bubo virginianus of the same country. It is doubtful whether some very light-colored adult males, supposed to belong to columbarius, as restricted, should not in reality be referred to this race, as the adult plumage of the male. But having seen no adult males from the region inhabited by the present bird obtained in the breeding-season, I am still in doubt whether the present form ever assumes the blue plumage.

As regards the climatic or regional modifications experienced by the Falco lithofalco on the American continent, the following summary of facts expresses my present views upon the subject. First: examples identical in all respects, or at least presenting no variations beyond those of an individual character, may be found from very widely separated localities; but the theory of explanation is, that individuals of one race may become scattered during their migrations, or wander off from their breeding-places. Second: the Atlantic region, the region of the plains, and the region of the northwest coast, have each a peculiar race, characterized by features which are also distinctive of races of other birds of the same region, namely, very dark—the dark tints intensified, and their area extended—in the northwest coast region; very light—the light markings extended and multiplied—in the middle region; and intermediate in the Atlantic region.

Habits. The distribution of the well-known Pigeon Hawk is very nearly coextensive with the whole of North America. It is found in the breeding-season as far to the north as Fort Anderson, on the Anderson and McKenzie rivers, ranging even to the Arctic coast. Specimens were taken by Mr. Ross at Lapierre House and at Fort Good Hope. Several specimens were taken by Mr. Dall at Nulato, where, he states, it is found all the year round. They were also taken by Bischoff at Kodiak. During the breeding-season it is found as far south as Nova Scotia, New Brunswick, and the northern portions of Maine, and probably Vermont and New York. It is abundant on the Pacific coast.