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4.5 Predicting breeding value
ОглавлениеBreeding value (or genetic merit) of an individual as a genetic parent is the sum of gene effects of the individual as measured by the performance of its progeny. Statistically, it is measured as twice the deviation of the offspring from the population mean (since the individual only contributes half of the alleles to its offspring). This estimate measures the ability of an individual to produce superior offspring. This is the part of an individual's genotypic value that is due to independent gene effects and hence can be transmitted. The mean breeding value becomes zero with random mating. This estimate is of importance to breeders because it assists them in selecting the best parents to use in their programs.
The Best Linear Unbiased Prediction(BLUP) is a common statistical method for estimating breeding values. It is unbiased because as more data are accumulated, the predicted breeding values approach the true values. BLUP is a method of estimating random effects. The context of this statistical method is the linear model
where y = is a vector of n observable random variables; B is vector of p unknown parameters with fixed value or effects; X and Z are known matrices; u and e are vectors of q and n, respectively, unobservable random variables (random effects).
To apply this technique, numerical scores are assigned to traits and compiled as predictions of the future. Simple traits can be most accurately and objectively measured and possibly predicted. Only one trait may be predicted in a model. This trait has to be objectively measurable with high accuracy. Further, it has to be heritable.