Читать книгу Migration of Birds (1979) - Frederick Charles Lincoln - Страница 18
Time of Day
ОглавлениеBecause most birds appear to be creatures of daylight, it seems remarkable that many should select the night for extended travel. Among the many nocturnal migrants are the smaller birds such as rails, flycatchers, orioles, most of the sparrows, the warblers, vireos, thrushes, and shorebirds. It is common to find woods and fields on one day almost barren of bird life and on the following day filled with sparrows, warblers, and thrushes, which indicates the arrival of migrants during the night. Waterfowl hunters sitting in their "blinds" frequently observe the passage of flocks of ducks and geese, but great numbers of these birds also pass through at night; the calls of Canada geese or the conversational gabbling of a flock of ducks are common night sounds in spring and fall in many parts of the country. Observations made with telescopes focused on the full moon have shown processions of birds, and one observer estimated their passage over his area at the rate of 9,000 per hour. This gives some indication of the numbers of birds in the air at night during peaks of migration. At such times radar observations have shown that nocturnal migration begins about an hour after sundown, reaches a peak shortly before midnight, and then gradually tapers off until daybreak. Unless special circuits are installed in radar sets, bird echoes during peak migration periods may cover a radar screen.
It has been suggested that small birds migrate by night to avoid their enemies. To a certain extent this may be true because the group includes not only weak fliers, such as the rails, but also the small song and insectivorous birds, such as wrens, small woodland flycatchers, and other species that habitually live more or less in concealment. These birds are probably much safer making their flights under the protecting cloak of darkness. Nevertheless, it must be remembered that night migrants include also the snipe, sandpipers, and plovers. Most shorebirds are usually found in the open and are among the more powerful fliers, as some of them make annual migratory flights over 2,000 miles nonstop across the ocean.
Night travel is probably best for the majority of birds chiefly from the standpoint of feeding. Digestion is very rapid in birds and yet the stomach of one killed during the day almost always contains food. To replace the energy required for long flight, it is essential that either food be obtained at comparatively short intervals or stores of fat be laid on prior to migration. If the smaller migrants were to make protracted flights by day they would arrive at their destination at nightfall almost exhausted, but since they are entirely daylight feeders, they would be unable to obtain food until the following morning. Unless reserve energy was carried in the form of fat, the inability to feed would delay further flights and result in great exhaustion or possibly even death should their evening arrival coincide with cold or stormy weather. By traveling at night, they can pause at daybreak and devote the entire period of daylight to alternate feeding and resting. This schedule permits complete recuperation and resumption of the journey on a subsequent evening after sufficient energy has been restored.
The day migrants include, in addition to some of the ducks and geese, the loons, cranes, gulls, pelicans, hawks, swallows, nighthawks, and swifts. Soaring birds, including broad-winged hawks, storks, and vultures, can only migrate during the day because their mode of flight makes them dependent on up-drafts created by heat from the sun for their long distance travels. On the other hand, swifts and swallows feed entirely on diurnal flying insects. The circling flocks are frequently seen in late summer feeding as they travel while working gradually southward. Formerly, great flocks of red-tailed, Swainson's, and rough-legged hawks could be seen wheeling majestically across the sky in the Plains States. In the East, good flights of broad-winged, Cooper's, and sharp-shinned hawks are still often seen, particularly along the Appalachian ridges.
Because many species of wading and swimming birds are able to feed at all hours, they migrate either by day or night and are not accustomed to seek safety in concealment. Some diving birds, including ducks that submerge when in danger, often travel over water by day and over land at night. Strong fliers like the snow geese can make the entire trip from their staging area in James Bay, Canada, to the wintering grounds on the Louisiana Gulf coast in one continuous flight. These birds are seldom shot by hunters en route between these two points but are often observed, when migrating, by aircraft pilots. Graham Cooch of the Canadian Wildlife Service tracked a flight of the blue phase of this species in 1955. The birds left James Bay on October 17 and arrived on the Gulf coast 60 hours later after an apparent continuous flight over the 1,700-mile route at an average speed of 28 miles per hour. Golden plovers, likewise, probably make the southward flight from the Arctic to the South American coast in one giant leap. Other Arctic species on their northward flight in the spring might prefer to fly at night in lower altitudes, but must necessarily fly during the day at higher altitudes because of the length of the days. Many warblers that normally fly at night may find themselves over water at daybreak and be forced to keep flying during the day until landfall is made.
Figure 3. Migration of the blackpoll warbler. As the birds move northward, the isochronal lines become farther apart, which indicates that the warblers move faster with the advance of spring. From April 30 to May 1 the average speed is about 30 miles per day, while from May 25 to May 30 it increases to more than 200 miles.
An interesting comparison of the flights of day and night migrants may be made through a consideration of the spring migrations of the blackpoll warbler and the cliff swallow. Both spend the winter as neighbors in South America, but when the impulse comes to start northward toward their respective breeding grounds, the warblers strike straight across the Caribbean Sea to Florida (Fig. 3.), while the swallows begin their journey by a westward flight of several hundred miles to Panama (Fig. 4.). From there they move leisurely along the western shore of the Caribbean Sea to Mexico, and, continuing to avoid a long trip over water, go completely around the western end of the Gulf of Mexico. This circuitous route adds more than 2,000 miles to the journey of the swallows that nest in Nova Scotia. The question may be asked: "Why should the swallow select a route so much longer and more roundabout than that taken by the blackpoll warbler?" The explanation is simple. The swallow is a day migrant while the warbler travels at night. The migration of the warbler is made up of a series of long nocturnal flights alternated with days of rest and feeding in favorable localities. The swallow, on the other hand, starts its migration several weeks earlier and catches each day's ration of flying insects during its aerial evolutions, while slowly migrating. The 2,000 extra miles flown along the insect-teeming shores of the Gulf of Mexico are exceeded by the great distances covered by these birds in normal pursuit of food.
Figure 4. Migration of the cliff swallow. A day migrant that, instead of flying across the Caribbean Sea as does the blackpoll warbler (see Fig. 3.), follows the coast of Central America, where food is readily obtained.
Although most of our smaller birds make their longest flights at night, close observation shows travel is continued to some extent by day. During the latter half of a migratory season birds may show evidence of an overpowering desire to hasten to their breeding grounds. At this time flocks of birds maintain a movement in the general direction of the seasonal journey while feeding on or near the ground. Sometimes they travel hurriedly, and while their flights may be short, they can cover an appreciable distance in the course of a day.