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The header is the most database-specific part of the record. Here, we will use the ENA version of the record for discussion (shown in its entirety in Appendix 1.1), with the corresponding DDBJ and GenBank versions of the header appearing in Appendix 1.2. The first line of the record provides basic identifying information about the sequence contained in the record, appropriately named the ID line; this corresponds to the LOCUS line in DDBJ/GenBank.

ID U54469; SV 1; linear; genomic DNA; STD; INV; 2881 BP.

The accession number is shown on the ID line, followed by its sequence version (here, the first version, or SV 1). As this is SV 1, this is equivalent to writing U54469.1, as described above. This is then followed by the topology of the DNA molecule (linear) and the molecule type (genomic DNA). The next element represents the ENA data class for this sequence (STD, denoting a “standard” annotated and assembled sequence). Data classes are used to group sequence records within functional divisions, enabling users to query specific subsets of the database. A description of these functional divisions can be found in Box 1.1. Finally, the ID line presents the taxonomic division for the sequence of interest (INV, for invertebrate; see Internet Resources) and its length (2881 base pairs). The accession number will also be shown separately on the AC line that immediately follows the ID lines.