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During spermiation, mature spermatids are released from SC into ST lumen prior to their subsequent passage to the epididymis. For spermiation, mature spermatids are oriented toward the lumen of the ST and subsequently released into the lumen by spermiation machinery over several days. Spermiation involves remodeling of the spermatid head and cytoplasm, removal of specialized adhesion structures between SC and mature spermatids, and the final expulsion of mature spermatid heads from the SC crypts [49].