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Char. Rictal bristles wanting; the loral feathers with bristly points; the frontal feathers generally not reaching to nostrils. Nostrils varied, exposed or not covered by feathers, and generally overhung by a scale-like membrane. Bill usually without notch (except in some Middle American genera). Wings much rounded, about equal to tail, which is graduated. Primaries ten, the first generally about half the second. Basal joint of middle toe usually united to half the basal joint of inner, and the whole of that of the outer, or more. Lateral toes about equal, or the outer a little the longer. Tarsi scutellate.

The impossibility of defining any large group of animals, so as to separate it stringently and abruptly from all others, is well understood among naturalists; and the Troglodytidæ form no exception to the rule. Some bear so close a resemblance to the Mocking Thrushes as to have been combined with them; while others again exhibit a close approximation to other subfamilies. The general affinities of the family, however, appear to be to the Turdidæ, and one of the best characters for separating the two families appears to exist in the structure of the feet.

In the Turdidæ the basal joint of the outer lateral toe is united to the middle toe, sometimes only a part of it; and the inner toe is cleft almost to its very base, so as to be opposable to the hind toe, separate from the others. In the Troglodytidæ, on the contrary, the inner toe is united by half its basal joint to the middle toe, sometimes by the whole of this joint; and the second joint of the outer toe enters wholly or partially into this union, instead of the basal joint only. In addition to this character, the open, exposed nostrils, the usually lengthened bill, the generally equal lateral toes, the short rounded wings, the graduated tail, etc., furnish points of distinction.

Genera.

A. Lateral toes very unequal.

a. Culmen depressed basally, the interval between the nostrils wider than the much compressed anterior half of the bill. Plate on the posterior half of the tarsus continuous. Catherpes.

b. Culmen compressed basally, the interval between the nostrils narrower than the rather depressed anterior half of the bill. Plate on the posterior half of the tarsus broken into smaller scales. Salpinctes.

B. Lateral toes equal.

c. Length about 8 inches. Campylorhynchus.

d. Length less than 6 inches.

Bill abruptly decurved or hooked at the tip. Outstretched feet not reaching near to end of tail. Thryothorus.

Tail longer than the wing, the feathers black, variegated terminally with whitish … Subgenus Thryomanes.

Tail shorter than the wing, the feathers rusty, not variegated with whitish … Subgenus Thryothorus.

Bill only gently curved at the tip. Outstretched feet reaching nearly to or beyond the end of the tail.

Back without streaks. No distinct superciliary stripe. Troglodytes.

Bill curved, sub-conical. Tail as long as wing … Subgenus Troglodytes.

Bill straight, subulate. Tail much shorter than wing … Subgenus Anorthura.

Back streaked with black and white. Cistothorus.

Bill short, stout; its depth equal to one half its length from the nostril; gonys straight or even convex, ascending. Crown streaked; no distinct superciliary stripe … Subgenus Cistothorus.

Bill elongated, slender; its depth less than one third its length from the nostril; gonys slightly concave, declining. Crown not streaked; a conspicuous superciliary stripe … Subgenus Telmatodytes.

Genus CAMPYLORHYNCHUS, Spix

Campylorhynchus, Spix, Av. Bras. I, 1824, 77. (Type, C. scolopaceus, Spix = Turdus variegatus, Gmel.)

Campylorhynchus brunneicapillus.

7149

Gen. Char. Bill stout, compressed, as long as, or longer than the head, without notch or rictal bristles; culmen and commissure curved; gonys nearly straight. Nostrils in the antero-inferior part of nasal groove, in advance of the frontal feathers, with an overhanging scale with thickened edge, as in Thryothorus; sometimes, as in the type, reduced to a slight ridge along the upper side of the nasal groove. Lateral septum not projecting below or anteriorly into the nasal cavity, but concealed by the nasal scale. Tarsus a little longer than middle toe and claw; claws strong, much curved, and very sharp; middle toe with basal joint adherent almost throughout. Wings and tail about equal, the latter graduated; the exterior webs of lateral feathers broad.

This genus embraces the largest species of the family, and is well represented in Middle and South America, two species only reaching into North America, which may be distinguished as follows:—

Top of head and post-ocular stripe reddish-brown; back streaked longitudinally and linearly with white. All the feathers beneath conspicuously spotted. Crissum and flanks with rounded or elongated spots. Iris reddish. Nostrils inferior, linear, overhung by a scale. Nests large and purse-shaped; eggs white, profusely marked with salmon-colored or reddish spots.

a. Spots much larger on throat and jugulum than elsewhere. Inner webs of second to fifth tail-feathers (between middle and outer feathers) black, except at tips. Length, 8.00; wing, 3.40; tail, 3.55. Hab. Adjacent borders of United States and Mexico … brunneicapillus.

b. Spots on throat and jugulum little larger than elsewhere. Inner webs of intermediate tail-feathers banded with white like the outer. Length, 7.50. Hab. Cape St. Lucas … affinis.

Campylorhynchus brunneicapillus, Gray

CACTUS WREN

Picolaptes brunneicapillus, Lafresnaye, Mag. de Zool. 1835, 61, pl. xlvii.—Lawr. Ann. N. Y. Lyc. V, 1851, 114.—Cassin, Birds Cal. Tex. 1854, 156, pl. xxv.—Heermann, J. A. N. Sc. II, 1853, 263. C. brunneicapillus, Gray, Genera, I, 1847, 159.—Bp. Consp. 1850, 223.—Scl. P. A. N. S. 156, 264.—Baird, Birds N. Am. 1858, 355; Pr. Phil. Acad. 1859, 3, etc.; Rev. 99.—Heermann, P. R. R. X, 1859.—Dresser, Ibis, 1865, 482 (Texas).—Cooper, Orn. Cal. 1, 1870, 61.

Sp. Char. Bill as long as the head. Above brown; darkest on the head, which is unspotted. Feathers on the back streaked centrally with white. Beneath whitish, tinged with rusty on the belly; the feathers of the throat and upper parts, and under tail-coverts, with large rounded black spots; those of the remaining under parts with smaller, more linear ones. Chin and line over the eye white. Tail-feathers black beneath, barred subterminally (the outer one throughout) with white. Iris, reddish-yellow. Length, 8 inches; wing, 3.40; tail, 3.55.

Hab. Adjacent borders of the United States and Mexico, from the mouth of the Rio Grande to the Valley of the Colorado, and to the Pacific coast of Southern California. Replaced at Cape St. Lucas by C. affinis.

This species is found abundantly along the line of the Rio Grande and Gila, extending northward some distance, and everywhere conspicuous by its wren-like habits and enormous nest.

Campylorhynchus brunneicapillus.

Habits. The Brown-headed Creeper is a comparatively recent addition to the fauna of the United States, but appears to be common along the southwestern borders of the United States, from the valley of the Rio Grande to San Diego, in California. In Lower California it is replaced by the C. affinis.

It was first added to our avifauna by Mr. Lawrence in 1851, on the strength of a specimen obtained in Texas by Captain McCown.

Dr. Heermann, in his paper on the Birds of California, speaks of finding it in the arid country back of Guymas, on the Gulf of California. This country, presenting only broken surfaces and a confused mass of volcanic rocks, covered by a scanty vegetation of thorny bushes and cacti, among other interesting birds, was found to contain this species in abundance. He describes it as a lively, sprightly species, uttering, at intervals, clear, loud, ringing notes. Its nest, composed of grasses and lined with feathers, was in the shape of a long purse, enormous for the size of the bird, and laid flat between the forks or on the branches of a cactus. The entrance was a covered passage, varying from six to ten inches in length. The eggs, six in number, he described as being of a delicate salmon-color, very pale, and often so thickly speckled with ash and darker salmon-colored spots as to give quite a rich cast to the whole surface of the egg.

Lieutenant Couch met with these birds near Monterey. He states that they have a rich, powerful song. Of the nest he gives substantially the same description as that furnished by Dr. Heermann.

The eggs are of an oblong-oval shape, slightly more pointed at one end, and are so equally and generally covered, over a white ground, with fine salmon-colored spots, as to present a uniform and almost homogeneous appearance. They vary in length from an inch to 1.02 inches, and have an average breadth of .68 of an inch.

Campylorhynchus affinis, Xantus

THE CAPE CACTUS WREN

Campylorhynchus affinis, Xantus, Pr. A. N. Sc. 1859, 298 (Cape St. Lucas).—Baird, Pr. A. N. Sc. 1859, 303; Rev. 100.—Scl. Catal. 1861, 17, No. 108.—Elliot, Illust. B. N. A. I, IV.—Cooper, Orn. Cal. 1, 1870, 62.

Sp. Char. Cap of head reddish-brown; the concealed centres of feathers dusky. Rest of upper parts grayish-brown, all the feathers of body and scapulars with broad central or shaft streaks of whitish edged with black; the streaks irregular in outline, on some feathers nearly linear, in others widening at intervals along the shaft. Outer webs of the wing-feathers crossed by about seven rows of whitish semicircular spots, with corresponding series of more circular ones on the inner web. Tail-feathers black, all of them with a series of about eight quadrate white spots on each web, which are alternate to each other, not opposite, and extend from or near the black shaft to the edge; the extreme tips of the feathers black; the two central feathers, however, more like the back, with irregular mottling of grayish and black. Upper tail-coverts barred transversely with black.

Under parts white, faintly tinged with rusty posteriorly; each feather spotted with black, excepting on the immaculate chin. These spots are rather larger and more quadrate on the jugulum, where they are sometimes on the sides of the feathers (on one or both sides); posteriorly, however, they are elongated or tear-shaped, and strung along the shaft, one or two on each. On the crissum they are large and much rounded, three or four on each longer feather. Legs rather dusky. Bill lead-color, pale at the base below; iris reddish-brown. A broad white stripe from bill over the eye and nape; edged above and below with black; line behind the eye like the crown; cheek-feathers white, edged with blackish.

Immature specimens exhibit a tendency to a whitish spotting in the ends of the feathers of the cap. A very young bird does not, however, differ materially, except in having the spots less distinct beneath, the white streaks less conspicuous above, the white of the wings soiled with rufous. Specimens vary considerably in the proportional as well as absolute thickness and length of the bill; thus, No. 32,167 measures .80 from nostril to end of bill, instead of .60, as given below for No. 12,965.

12,965. Total length, 7.50; wing, 3.30; tail, 3.40; its graduation, .45; exposed portion of first primary, 1.42, of second, 2.15, of longest, or fourth (measured from exposed base of first primary), 2.45; length of bill from forehead, .90, from nostril, .60; along gape, 1.07; tarsus, 1.02; middle toe and claw, .90; claw alone, .25; hind toe and claw, .76; claw alone, .35.

Hab. Only observed at Cape St. Lucas, Lower California.

This species is most nearly allied to C. brunneicapillus; the most apparent difference at first sight being in the greater concentration of black on the throat and jugulum in brunneicapillus, and the much smaller size of the remaining spots on the under parts, with the decided light-cinnamon of the posterior portion of the body. The outer and central tail-feathers alone are marked as in C. affinis, the intermediate ones being entirely black, with the exception of a white subterminal band.

This is one of the most characteristic birds constituting the isolated fauna of Cape St. Lucas. Like nearly all the species peculiar to this remarkable locality, it is exceedingly abundant, breeding in immense numbers. It has not yet been detected elsewhere, though it may possibly be found on the Lower Colorado.

Habits. This recently described species was first discovered by Mr. Xantus, and has, so far as is known, a somewhat restricted locality, having been met with only at the southern extremity of Lower California, where it is an exceedingly abundant bird. Mr. Xantus has published no observations in regard to its habits, which, however, are probably very nearly identical with those of the more common species. From the brief memoranda given by him in the general register of his collections, made at Cape St. Lucas, we gather that their nests were built almost exclusively in opuntias, cacti, and the prickly pear, and were generally only four or five feet from the ground, but occasionally at the height of ten feet.

The nests are large purse-shaped collections of twigs and coarse grasses, very similar to, and hardly distinguishable in any respect from, those of the more northern species. The eggs vary from 1.05 to 1 inch in length, and from .65 to .70 of an inch in breadth, and have a reddish-white ground very uniformly dotted with fine markings of reddish-brown, purple, and slate.

Subgenus SALPINCTES, Cabanis

Salpinctes, Cabanis, Wiegmann’s Archiv, 1847, I, 323. (Type, Troglodytes obsoletus, Say.)

Gen. Char. Bill as long as the head; all the outlines nearly straight to the tip, then decurved; nostrils oval. Feet weak; tarsi decidedly longer than the middle toe; outer lateral toe much longer, reaching to the base of the middle claw, and equal to the hinder. Wings about one fifth longer than the tail; the exposed portion of the first primary about half that of the second, and two fifths that of the fourth and fifth. Tail-feathers very broad, plane, nearly even or slightly rounded; the lateral moderately graduated.

Of this genus but one species is so far known in the United States, the Rock Wren of the earlier ornithologists. It is peculiar among its cognate genera by having the two continuous plates on each side the tarsus divided into seven or more smaller plates, with a naked interval between them and the anterior scutellæ. Other characters will be found detailed in the Review of American Birds, p. 109.

Salpinctes obsoletus, Caban

ROCK WREN

Troglodytes obsoletus, Say, Long’s Exped. II, 1823, 4 (south fork of Platte).—Aud. Orn. Biog. IV, pl. ccclx.—Ib. B. A. II, pl. cxvi.—Newberry, P. R. R. Rep. VI, IV, 1857, 80.—Heermann, P. R. R. Rep. X, 1859, 41. Salpinctes obsoletus, Cab. Wiegmann’s Archiv, 1847, I, 323.—Baird, Birds N. Am. 1858, 357; Rev. 110.—Sclater, P. Z. S. 1859, 371 (Oaxaca).—Cooper, Orn. Cal. 1, 1870, 64. ? Troglodytes latisfasciatus, Licht. Preis-Verzeich. 1831, No. 82.

Salpinctes obsoletus.

7157 ♂

Sp. Char. Plumage very soft and lax. Bill about as long as the head. Upper parts brownish-gray, each feather with a central line and (except on the head) transverse bars of dusky, and a small dull brownish-white spot at the end (seen also on the tips of the secondaries). Rump, sides of the body, and posterior part of belly and under tail-coverts dull cinnamon, darker above. Rest of under parts dirty white; feathers of throat and breast with dusky central streaks. Lower tail-coverts banded broadly with black. Inner tail-feathers like the back, the others with a broad black bar near the end; the tips cinnamon; the outer on each side alternately banded with this color and black. A dull white line above and behind the eye. Iris brown. Length, 5.70; wing, 2.82; tail, 2.40. Young not marked or banded beneath. Eggs white, spotted with red.