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NORTH AMERICAN BIRDS
Family TURDIDÆ.—The Thrushes
ОглавлениеThe Turdidæ, with the Saxicolidæ and Cinclidæ, form a group closely related, by common characters, and appreciably different from the other Oscines with slender bills and specially insectivorous habits, having, like them, ten primaries (the first much shorter than the second, but nearly always appreciable), and the nostrils uncovered. The great family of Sylvicolidæ, with similar characters of the bill, never present more than nine primaries. The most striking of these common characters is seen in the deeply cleft toes, of which the outer is united by the basal joint alone to the middle toe, while the inner is separated almost to the very base of its first joint.21 The frontal feathers extend, with rare exceptions, to the very nostrils. The bill is elongated and subulate, moderately slender, and usually notched at tip; the culmen moderately curved from the base, and the mouth well provided with bristles, except in a few cases. Usually the scutellæ covering the front and sides of the tarsus are fused into one continuous plate, or else scarcely appreciable, except on the inner edge only; in the Mocking Thrushes they are, however, distinctly marked. The lateral toes are nearly equal, the outer rather the longer. With these as some of the principal characteristics, they may be distinguished from each other as follows:—
Note.—In the present work the length of the tail is measured from the coccyx, inside of the skin, and not, as usually the case, from the base of the quills at their insertion. The wings are measured from the carpal joint, with dividers.
A. Nostrils oval. Loral and frontal feathers with bristly points, or interspersed with bristles; rictus with longer or shorter bristles.
Saxicolidæ. Wings very long and much pointed, reaching beyond the middle of the short square or emarginated tail, and one and a half times or more the length of the latter. The spurious primary very short, the second quill longer than the fourth. In the closed wing the outer secondary reaches only about two thirds the length of longest primary.
Turdidæ. Wings moderate, more rounded, not reaching beyond middle of the often rounded tail, and not more than one and a third the latter, usually more nearly equal. Spurious primary sometimes half the length of second quill; the second quill shorter than the fourth. In the closed wing the outer secondary reaches three fourths or more the length of longest primary.
B. Nostrils linear, in lower edge of nasal membrane. Loral and frontal feathers soft and downy, and no bristles or bristly points whatever about the mouth.
Cinclidæ. Body very short and broad. Wings short, rounded, and concave.
The American Sylviidæ are in some respects very closely related to the Saxicolidæ, but may be distinguished by their much smaller size, more slender and depressed bill, more strongly bristled rictus, etc.; on which account they are more strictly “fly-catchers,” taking their prey in great part on the wing.
Of the three families, the Turdidæ contain a great variety of forms, and exhibit widely different characters, rendering it exceedingly difficult to arrange them in any systematic or regular sequence, or to accurately define their boundaries. In the Birds of North America, the Mocking Thrushes were placed among the Wrens, on account of the distinct tarsal scutellæ, and other characters. We are now, however, inclined to believe, with Dr. Sclater, that their place is with the recognized Turdidæ; and, among other reasons, on the ground of their more deeply cleft toes, and greater extension forward of frontal feathers. The following synopsis of the North American forms will serve the purpose of determining the genera, even if these are not arranged or combined in a strictly natural manner.
A. Turdinæ.—Tarsus covered anteriorly with a continuous plate without scales.
Wings decidedly longer than the tail, which is nearly even. Bill considerably shorter than the head.
First quill usually not one fourth the second. Wings pointed. Tarsus hardly the length of head, but yet longer than middle toe; outstretched toes falling short of tip of tail … Turdus.
B. Miminæ.—Tarsi scutellate anteriorly; scales seven.
Wings decidedly longer than the tail, which is nearly even. Tarsus as long as the head.
Bill decidedly shorter than the head, scarcely notched; wings pointed; first quill less than half the second, third and fourth longest. Claws not peculiar. Bristles prominent. Tarsus considerably longer than middle toe and claw … Oreoscoptes.
Wings decidedly shorter than the tail, which is considerably graduated; first quill half or more than half the second.
Bill notched at tip, shorter than head; straight.
Scutellæ very distinct; gonys straight, or even declining at tip … Mimus.
Scutellæ more or less obsolete; gonys convex, ascending at tip … Galeoscoptes.
Bill not notched at tip, lengthened; sometimes much decurved … Harporhynchus.
Note.—In the Review of American Birds, I., May, 1866, 409, I have advanced the suggestion that the N. American genus Myiadestes, usually placed under the Ampelidæ, really belongs under Turdidæ in a group Myiadestinæ. The relationships are certainly very close, as is shown by the characters given below.
Common Characters.—Tarsi without regular transverse scutellæ, except at lower end. Wings acute, pointed, as long as or longer than tail, which is but slightly graduated. First primary rarely half second, which exceeds the secondaries. Base of quills buffy yellow, as are inner edges. Tail spotted or varied at the end. Young birds with many light spots. Very melodious singers.
Myiadestinæ. Bill short, much depressed; mouth deeply cleft; width at base about equal to the distance from nostril to tip, or greater; commissure more than twice distance from nostrils to tip of bill, and nearly two and a half times length of gonys. Legs weak; tarsi rather longer than middle toe and claw. Tail feathers tapering slightly from base to near tip, giving a slightly cuneate appearance to the tail.
Turdinæ. Bill stouter, more lengthened; narrow at base and more compressed; width at base less than distance from nostril to tip; commissure not more than twice distance from nostrils to tip of bill, and about twice length of gonys. Tarsi stouter, longer than middle toe and claw. Tail feathers widening slightly from base to near tip, giving a parallel-sided or slightly fan-shaped appearance to the tail.
The Miminæ differ, as already mentioned, in the scutellate tarsi: more rounded wings, etc.—S. F. B.
21
In a perfectly fresh specimen of Turdus mustelinus, the basal half of the first phalanx of the inner toe is connected with the first joint of the middle toe by a membrane which stretches across to within two fifths of the end of the latter; there appears, however, to be no ligamentous adhesion. The basal joint of the outer toe is entirely adherent, and a membrane extends from nearly the basal half of the second joint to the distal end of the first joint of the middle toe. When this connecting membrane becomes dried the division of the toes appears considerably greater.
When the toes are all extended in line with the tarsus, the hind claw stretches a little beyond the lateral and scarcely reaches the base of the middle claw.
The plates at the upper surface of the basal joints of the toes are quadrangular and opposite each other.