The Origin of Species

The Origin of Species
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HarperCollins is proud to present its new range of best-loved, essential classics.‘How fleeting are the wishes and efforts of man! how short his time! and consequently how poor will his products be, compared with those accumulated by nature during whole geological periods.’Still considered one of the most important and groundbreaking works of science ever written, Darwin’s eminently readable exploration of the evolutionary process challenged most of the strong beliefs of the Western world. Forced to question the idea of the Creator, mid-nineteenth century readers were faced with Darwin’s theories on the laws of natural selection and the randomness of evolution, causing massive controversy at the time. However, Darwin’s theories remain instrumental in providing the backbone to modern biology today.

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Charles Darwin. The Origin of Species

THE. ORIGIN OF. SPECIES. Charles Darwin

CONTENTS

INTRODUCTION

CHAPTER 1 Variation Under Domestication

Causes of Variability

Effects of Habit and of The Use or Disuse of Parts; Correlated Variation; Inheritance

Character of Domestic Varieties; Difficulty of Distinguishing Between Varieties and Species; Origin of Domestic Varieties From One or More Species

Breeds of The Domestic Pigeon, Their Differences and Origin

Principles of Selection Anciently Followed, and Their Effects

Unconscious Selection

Circumstances Favourable to Man’s Power of Selection

CHAPTER 2 Variation Under Nature

Individual Differences

Doubtful Species

Wide-Ranging, Much Diffused, and Common Species Vary Most

Species of The Larger Genera in Each Country Vary More Frequently Than The Species of The Smaller Genera

Many of The Species Included Within The Larger Genera Resemble Varieties in Being Very Closely, but Unequally, Related to Each other, and in having Restricted Ranges

Summary

CHAPTER 3 Struggle for Existence

The Term, Struggle for Existence, Used in a Large Sense

Geometrical Ratio of Increase

Nature of The Checks to Increase

Complex Relations of All Animals and Plants to Each Other in The Struggle for Existence

Struggle for Life Most Severe between Individuals and Varieties of The Same Species

CHAPTER 4 Natural Selection; or the Survival of the Fittest

Sexual Selection

Illustrations of the Action of Natural Selection, or the Survival of the Fittest

Circumstances Favourable for The Production of New Forms Through Natural Selection

Extinction Caused by Natural Selection

Divergence of Character

The Probable Effects of the Action of Natural Selection Through Divergence of Character and Extinction, on the Descendants of a Common Ancestor

On The Degree to which Organisation Tends to Advance

Convergence of Character

Summary of Chapter

CHAPTER 5 Laws of Variation

Effects of the Increased Use and Disuse of Parts, as Controlled by Natural Selection

Acclimatisation

Correlated Variation

Compensation and Economy of Growth

Multiple, Rudimentary, and Lowly-Organised Structures are Variable

A Part Developed in any Species in an Extraordinary Degree or Manner, in Comparison with the Same Part in Allied Species, Tends to be Highly Variable

Specific Characters More Variable than Generic Characters

Secondary Sexual Characters Variable

Distinct Species Present Analogous Variations, So that a Variety of One Species often Assumes a Character Proper to an Allied Species, or Reverts to Some of the Characters of an Early Progenitor

CHAPTER 6 Difficulties of The Theory

On the Absence or Rarity of Transitional Varieties

On the Origin and Transition of Organic beings with Peculiar Habits and Structure

Organs of Extreme Perfection and Complication

Modes of Transition

Special Difficulties of the Theory of Natural Selection

Organs of Little Apparent Importance, as affected by Natural Selection

Utilitarian Doctrine, How Far True: Beauty, How Acquired

Summary: The Law of Unity of Type and of the Conditions of Existence Embraced by The Theory of Natural Selection

CHAPTER 7 Miscellaneous Objections to The Theory of Natural Selection

CHAPTER 8 Instinct

Inherited Changes of Habit or Instinct in Domesticated Animals

Special Instincts

Instincts of the Cuckoo

Slave-Making Instinct

Cell-Making Instinct of The Hive-Bee

Objections to The Theory of Natural Selection as Applied to Instincts: Neuter and Sterile Insects

Summary

CHAPTER 9 Hybridism

Degrees of Sterility

Laws Governing The Sterility of First Crosses and of Hybrids

Origin and Causes of The Sterility of First Crosses and of Hybrids

Reciprocal Dimorphism and Trimorphism

Fertility of Varieties when Crossed, and of their Mongrel Offspring, not Universal

Hybrids and Mongrels Compared, Independently of Their Fertility

Summary of Chapter

CHAPTER 10 On the Imperfection of the Geological Record

On The Lapse of Time, as Inferred from The Rate of Deposition and Extent of Denudation

On the Poorness of Palaeontological Collections

On the Absence of Numerous Intermediate Varieties in any Single Formation

On The Sudden Appearance of Whole Groups of Allied Species

On the Sudden Appearance of Groups of Allied Species in the Lowest Known Fossiliferous Strata

CHAPTER 11 On The Geological Succession of Organic Beings

On Extinction

On the Forms of Life Changing Almost Simultaneously Throughout The World

On the Affinities of Extinct Species to Each Other, and to Living Forms

On the State of Development of Ancient Compared with Living Forms

On the Succession of the Same Types within the Same Areas, during the Later Tertiary Periods

Summary of the Preceding and Present Chapters

CHAPTER 12 Geographical Distribution

Single Centres of Supposed Creation

Means of Dispersal

Dispersal during the Glacial Period

Alternate Glacial Periods in The North and South

CHAPTER 13 Geographical Distribution—Continued

Fresh-water Productions

On the Inhabitants of Oceanic Islands

Absence of Batrachians and Terrestrial Mammals on Oceanic Islands

On the Relations of the Inhabitants of Islands to those of the Nearest Mainland

Summary of The Last and Present Chapters

CHAPTER 14 Mutual Affinities of Organic Beings

Classification

Analogical Resemblances

On the Nature of the Affinities Connecting Organic Beings

Morphology

Development and Embryology

Rudimentary, Atrophied, and Aborted Organs

Summary

CHAPTER 15 Recapitulation and Conclusion

GLOSSARY OF THE PRINCIPAL SCIENTIFIC TERMS USED IN THE PRESENT VOLUME

CLASSIC LITERATURE: WORDS AND PHRASES adapted from theCollins English Dictionary

ABOUT THE AUTHOR. Life & Times

Alfred Russell Wallace

The Discovery

HISTORY OF COLLINS

COPYRIGHT

ABOUT THE PUBLISHER

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Cover

Title Page

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I will now say a few words on the circumstances, favourable or the reverse, to man’s power of selection. A high degree of variability is obviously favourable, as freely giving the materials for selection to work on; not that mere individual differences are not amply sufficient, with extreme care, to allow of the accumulation of a large amount of modification in almost any desired direction. But as variations manifestly useful or pleasing to man appear only occasionally, the chance of their appearance will be much increased by a large number of individuals being kept. Hence number is of the highest importance for success. On this principle Marshall formerly remarked, with respect to the sheep of part of Yorkshire, “As they generally belong to poor people, and are mostly IN SMALL LOTS, they never can be improved.” On the other hand, nurserymen, from keeping large stocks of the same plant, are generally far more successful than amateurs in raising new and valuable varieties. A large number of individuals of an animal or plant can be reared only where the conditions for its propagation are favourable. When the individuals are scanty all will be allowed to breed, whatever their quality may be, and this will effectually prevent selection. But probably the most important element is that the animal or plant should be so highly valued by man, that the closest attention is paid to even the slightest deviations in its qualities or structure. Unless such attention be paid nothing can be effected. I have seen it gravely remarked, that it was most fortunate that the strawberry began to vary just when gardeners began to attend to this plant. No doubt the strawberry had always varied since it was cultivated, but the slight varieties had been neglected. As soon, however, as gardeners picked out individual plants with slightly larger, earlier, or better fruit, and raised seedlings from them, and again picked out the best seedlings and bred from them, then (with some aid by crossing distinct species) those many admirable varieties of the strawberry were raised which have appeared during the last half-century.

With animals, facility in preventing crosses is an important element in the formation of new races—at least, in a country which is already stocked with other races. In this respect enclosure of the land plays a part. Wandering savages or the inhabitants of open plains rarely possess more than one breed of the same species. Pigeons can be mated for life, and this is a great convenience to the fancier, for thus many races may be improved and kept true, though mingled in the same aviary; and this circumstance must have largely favoured the formation of new breeds. Pigeons, I may add, can be propagated in great numbers and at a very quick rate, and inferior birds may be freely rejected, as when killed they serve for food. On the other hand, cats, from their nocturnal rambling habits, can not be easily matched, and, although so much valued by women and children, we rarely see a distinct breed long kept up; such breeds as we do sometimes see are almost always imported from some other country. Although I do not doubt that some domestic animals vary less than others, yet the rarity or absence of distinct breeds of the cat, the donkey, peacock, goose, etc., may be attributed in main part to selection not having been brought into play: in cats, from the difficulty in pairing them; in donkeys, from only a few being kept by poor people, and little attention paid to their breeding; for recently in certain parts of Spain and of the United States this animal has been surprisingly modified and improved by careful selection; in peacocks, from not being very easily reared and a large stock not kept; in geese, from being valuable only for two purposes, food and feathers, and more especially from no pleasure having been felt in the display of distinct breeds; but the goose, under the conditions to which it is exposed when domesticated, seems to have a singularly inflexible organisation, though it has varied to a slight extent, as I have elsewhere described.

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