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Other efforts directed at improving the cellular microenvironment take into account that atmospheric oxygen conditions are not optimal for most mammalian cells in culture. Brackertz et al.⁶⁴⁶ and Held and Sönnichsen⁶⁴⁷ demonstrated improved AFC growth under hypoxic conditions. If multigas incubators are opened frequently, however, their humidity tends to decrease, which could disadvantage cells growing in open dishes.